PANTHER Gene Information

Gene Symbol(s):

DDX39A

Organism:

Bos taurus

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Tree

Gene Name:

RNA helicase

Gene ID:

ENSBTAG00000021820.3

Protein ID:

Q5E970

Persistent Id:

PTN002776429

Alternate Ids:

ENSBTAP00000068550(Ensembl_PRO)	77736449(GI)
Q5E970(Synonym)	ENSBTAT00000066612(Ensembl_TRS)
AAX09067(EMBL-CDS)	BT021050(EMBL)
614215(GeneID)	ENSBTAG00000021820(Ensembl)
Q5E970_BOVIN(UniProtKB-ID)	XM_015471952(refSeq)
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PANTHER Classification

PANTHER Family:	ATP-DEPENDENT RNA HELICASE DBP3 (PTHR47958)
PANTHER Subfamily:	ATP-DEPENDENT RNA HELICASE DDX39A (PTHR47958:SF10)
PANTHER GO-slim Molecular Function:	helicase activity mRNA binding catalytic activity, acting on RNA
PANTHER GO-slim Biological Process:	mRNA export from nucleus mRNA splicing, via spliceosome
PANTHER GO-slim Cellular Component:
PANTHER protein class:	RNA helicase

Pathway Categories:

No pathway information available

GENE ONTOLOGY DATABASE ANNOTATIONS

Click to Expand...

GO MF Complete:	ATP binding, identical protein binding, RNA helicase activity, mRNA binding, hydrolase activity
GO BP Complete:	mRNA export from nucleus, mRNA splicing, via spliceosome
GO CC Complete:	cytoplasm, nuclear speck

REACTOME DATABASE ANNOTATIONS Click to Expand...

Reactome Pathways:

mRNA 3'-end processing, Gene expression (Transcription), Metabolism of RNA, RNA Polymerase II Transcription, RNA Polymerase II Transcription Termination, Transport of Mature mRNA derived from an Intron-Containing Transcript, Processing of Capped Intron-Containing Pre-mRNA, Transport of Mature Transcript to Cytoplasm

Orthologs

ID	Organism	Type
HUMAN\|HGNC=17821\|UniProtKB=O00148	Homo sapiens	LDO
PANTR\|Ensembl=ENSPTRG00000042673.1\|UniProtKB=K7A9J4	Pan troglodytes	LDO
GORGO\|Ensembl=ENSGGOG00000005414.3\|UniProtKB=G3RN98	Gorilla gorilla gorilla	LDO
MOUSE\|MGI=MGI=1915528\|UniProtKB=Q8VDW0	Mus musculus	LDO
RAT\|RGD=619920\|UniProtKB=Q5U216	Rattus norvegicus	LDO
PIG\|Ensembl=ENSSSCG00000013776.5\|UniProtKB=A0A287BN69	Sus scrofa	LDO
HORSE\|Ensembl=ENSECAG00000040285.3\|UniProtKB=A0A3Q2HBP7	Equus caballus	LDO
FELCA\|Ensembl=ENSFCAG00000013488.6\|UniProtKB=A0A5F5Y5K0	Felis catus	LDO
CANLF\|Ensembl=ENSCAFG00845022719.1\|UniProtKB=A0A8I3PGD1	Canis lupus familiaris	LDO
MONDO\|Ensembl=ENSMODG00000002448.4\|UniProtKB=A0A5F8H7S6	Monodelphis domestica	LDO
ANOCA\|Ensembl=ENSACAG00000010756.4\|UniProtKB=G1KKJ7	Anolis carolinensis	LDO
XENLA\|Gene=ddx39a.L\|UniProtKB=Q7ZX82	Xenopus laevis	O
XENLA\|Gene=ddx39a.S\|UniProtKB=A0A8J1MNW5	Xenopus laevis	LDO
XENTR\|Xenbase=XB-GENE-966447\|UniProtKB=Q28EE7	Xenopus tropicalis	LDO
ORYLA\|Ensembl=ENSORLG00000000279.2\|UniProtKB=H2L3M0	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000010565.2\|UniProtKB=H2M484	Oryzias latipes	LDO
DANRE\|ZFIN=ZDB-GENE-030131-4275\|UniProtKB=A2AWA1	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-040426-2902\|UniProtKB=Q803W0	Danio rerio	LDO
LEPOC\|Ensembl=ENSLOCG00000006534.1\|UniProtKB=W5MHN7	lepisosteus oculatus	LDO
CIOIN\|Ensembl=ENSCING00000005175.3\|UniProtKB=F6Z194	Ciona intestinalis	LDO
BRAFL\|Gene=LOC118420743\|UniProtKB=Q8T783	Branchiostoma floridae	LDO
STRPU\|EnsemblGenome=LOC580001\|UniProtKB=A0A7M7LWK3	Strongylocentrotus purpuratus	LDO
CAEEL\|WormBase=WBGene00001840\|UniProtKB=Q18212	Caenorhabditis elegans	LDO
CAEBR\|WormBase=WBGene00024046\|UniProtKB=A8WNA1	Caenorhabditis briggsae	LDO
PRIPA\|EnsemblGenome=WBGene00112028\|UniProtKB=A0A2A6CFS0	Pristionchus pacificus	LDO
DROME\|FlyBase=FBgn0014189\|UniProtKB=Q27268	Drosophila melanogaster	LDO
ANOGA\|EnsemblGenome=AGAP009075\|UniProtKB=A0A1S4H010	Anopheles gambiae	LDO
TRICA\|EnsemblGenome=TC014413\|UniProtKB=D6WLU7	Tribolium castaneum	LDO
DAPPU\|EnsemblGenome=DAPPUDRAFT_309072\|UniProtKB=E9G3G8	Daphnia pulex	O
DAPPU\|EnsemblGenome=DAPPUDRAFT_219465\|UniProtKB=E9HTR5	Daphnia pulex	LDO
HELRO\|EnsemblGenome=HelroG112898\|UniProtKB=T1EFN1	helobdella robusta	O
HELRO\|EnsemblGenome=HelroG178681\|UniProtKB=T1FDK3	helobdella robusta	O
HELRO\|EnsemblGenome=HelroG83915\|UniProtKB=T1G5B8	helobdella robusta	O
HELRO\|EnsemblGenome=HelroG112849\|UniProtKB=T1EFM6	helobdella robusta	LDO
NEMVE\|EnsemblGenome=NEMVEDRAFT_v1g236766\|UniProtKB=A7RYW1	Nematostella vectensis	LDO
TRIAD\|EnsemblGenome=TriadG63649\|UniProtKB=B3RP53	Trichoplax adhaerens	LDO
MONBE\|EnsemblGenome=MONBRDRAFT_36918\|UniProtKB=A9UYD6	Monosiga brevicollis	LDO
YEAST\|SGD=S000002242\|UniProtKB=Q07478	Saccharomyces cerevisiae	LDO
EREGS\|EnsemblGenome=AGOS_ADR260C\|UniProtKB=Q759L6	Eremothecium gossypii	LDO
CANAL\|CGD=CAL0000200240\|UniProtKB=A0A1D8PKZ3	Candida albicans	LDO
YARLI\|EnsemblGenome=YALI0_A11157g\|UniProtKB=Q6CH90	Yarrowia lipolytica	LDO
EMENI\|Gene_ORFName=AN8722\|UniProtKB=Q5ASK8	Emericella nidulans	LDO
ASPFU\|EnsemblGenome=AFUA_6G02630\|UniProtKB=Q4WCW2	Neosartorya fumigata	LDO
NEUCR\|EnsemblGenome=NCU09076\|UniProtKB=Q7RW10	Neurospora crassa	LDO
SCLS1\|EnsemblGenome=SS1G_05270\|UniProtKB=A7EIX7	Sclerotinia sclerotiorum	LDO
PHANO\|EnsemblGenome=SNOG_15623\|UniProtKB=Q0TXZ2	Phaeosphaeria nodorum	LDO
SCHPO\|PomBase=SPAC17G6.14c\|UniProtKB=O13792	Schizosaccharomyces pombe	LDO
SCHJY\|EnsemblGenome=SJAG_04589\|UniProtKB=B6K783	Schizosaccharomyces japonicus	LDO
CRYNJ\|EnsemblGenome=CNA07200\|UniProtKB=P0CQ96	Cryptococcus neoformans	LDO
USTMA\|EnsemblGenome=UMAG_11769\|UniProtKB=A0A0D1BZF0	Ustilago maydis	LDO
PUCGT\|EnsemblGenome=PGTG_08191\|UniProtKB=E3KCJ4	Puccinia graminis	LDO
BATDJ\|Gene_ORFName=BATDEDRAFT_15696\|UniProtKB=F4NTX2	Batrachochytrium dendrobatidis	LDO
ENTH1\|EnsemblGenome=CL6EHI_036900\|UniProtKB=C4M9V7	Entamoeba histolytica	O
ENTH1\|EnsemblGenome=CL6EHI_151600\|UniProtKB=C4LSK1	Entamoeba histolytica	LDO
ENTH1\|Gene_ORFName=EHI_150160\|UniProtKB=B1N3F1	Entamoeba histolytica	LDO
DICDI\|dictyBase=DDB_G0269932\|UniProtKB=Q55CR6	Dictyostelium discoideum	LDO
DICPU\|EnsemblGenome=DICPUDRAFT_25293\|UniProtKB=F0Z6V0	Dictyostelium purpureum	LDO
LEIMA\|EnsemblGenome=LMJF_21_1552\|UniProtKB=Q4QC38	Leishmania major	LDO
TRYB2\|Gene_ORFName=Tb10.70.7730\|UniProtKB=Q38CI6	Trypanosoma brucei	LDO
GIAIC\|Gene_ORFName=GL50803_0016376\|UniProtKB=A0A644F7Y3	Giardia intestinalis	LDO
TRIV3\|Gene_ORFName=TVAG_179860\|UniProtKB=A2F468	Trichomonas vaginalis	LDO
PLAF7\|EnsemblGenome=PF3D7_0209800\|UniProtKB=Q9TY94	Plasmodium falciparum	LDO
PARTE\|EnsemblGenome=GSPATT00025153001\|UniProtKB=A0EAV2	Paramecium tetraurelia	LDO
PARTE\|EnsemblGenome=GSPATT00001774001\|UniProtKB=A0D6R6	Paramecium tetraurelia	LDO
PARTE\|EnsemblGenome=GSPATT00009104001\|UniProtKB=A0CPJ5	Paramecium tetraurelia	O
PARTE\|EnsemblGenome=GSPATT00005317001\|UniProtKB=A0BPV0	Paramecium tetraurelia	O
THAPS\|EnsemblGenome=THAPSDRAFT_269556\|UniProtKB=B8LBP2	Thalassiosira pseudonana	LDO
PHYRM\|Gene=H3G9Z6_PHYRM\|UniProtKB=H3G9Z6	Phytophthora ramorum	LDO
CHLRE\|EnsemblGenome=CHLRE_17g719000v5\|UniProtKB=A8JIJ1	Chlamydomonas reinhardtii	LDO
LACSA\|Gene_ORFName=LSAT_V11C400192480\|UniProtKB=A0A9R1XGF7	Lactuca sativa	LDO
PHYPA\|EnsemblGenome=Pp3c11_18810\|UniProtKB=A9SV98	Physcomitrella patens	LDO
AMBTC\|EnsemblGenome=AMTR_s00036p00225190\|UniProtKB=U5CZ48	Amborella trichopoda	LDO
ZOSMR\|Gene_ORFName=ZOSMA_387G00010\|UniProtKB=A0A0K9P767	Zostera marina	O
NELNU\|Gene_ORFName=HUJ06_018873\|UniProtKB=A0A1U8AHB2	Nelumbo nucifera	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a006871mg\|UniProtKB=A0A022QAL3	Erythranthe guttata	O
MANES\|EnsemblGenome=MANES_04G114500\|UniProtKB=A0A2C9W1L9	Manihot esculenta	O
MANES\|EnsemblGenome=MANES_11G054900\|UniProtKB=A0A2C9UZW9	Manihot esculenta	O
ZOSMR\|Gene_ORFName=ZOSMA_212G00090\|UniProtKB=A0A0K9PKK6	Zostera marina	LDO
MUSAM\|EnsemblGenome=Ma03_g27200\|UniProtKB=A0A804IGU2	Musa acuminata subsp. malaccensis	O
MUSAM\|EnsemblGenome=Ma08_g05230\|UniProtKB=A0A804K347	Musa acuminata subsp. malaccensis	LDO
SETIT\|EnsemblGenome=SETIT_001080mg\|UniProtKB=K3XGQ9	Setaria italica	O
SORBI\|EnsemblGenome=SORBI_3003G186900\|UniProtKB=A0A1B6Q469	Sorghum bicolor	O
MAIZE\|EnsemblGenome=Zm00001eb356560\|UniProtKB=C4IZ62	Zea mays	O
SETIT\|EnsemblGenome=SETIT_025481mg\|UniProtKB=K3ZG75	Setaria italica	LDO
SORBI\|EnsemblGenome=SORBI_3009G257600\|UniProtKB=A0A1B6PAQ6	Sorghum bicolor	LDO
MAIZE\|EnsemblGenome=Zm00001eb345420\|UniProtKB=K7UTZ2	Zea mays	LDO
ORYSJ\|EnsemblGenome=Os01g0550000\|UniProtKB=Q5JK84	Oryza sativa	LDO
ORYSJ\|EnsemblGenome=Os01g0549700\|UniProtKB=Q0JM17	Oryza sativa	LDO
BRADI\|EnsemblGenome=BRADI_2g40800v3\|UniProtKB=I1HNH1	Brachypodium distachyon	O
BRADI\|EnsemblGenome=BRADI_2g40780v3\|UniProtKB=I1HNG8	Brachypodium distachyon	O
BRADI\|EnsemblGenome=BRADI_2g40807v3\|UniProtKB=A0A0Q3GAC3	Brachypodium distachyon	O
HORVV\|EnsemblGenome=HORVU.MOREX.r3.3HG0264900\|UniProtKB=A0A8I6XYL6	Hordeum vulgare subsp. vulgare	O
WHEAT\|EnsemblGenome=TraesCS3A02G200100\|UniProtKB=A0A3B6EHA2	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS3D02G204400\|UniProtKB=A0A3B6GVX0	Triticum aestivum	O
HORVV\|EnsemblGenome=HORVU.MOREX.r3.1HG0095070\|UniProtKB=A0A8I6WZ89	Hordeum vulgare subsp. vulgare	LDO
WHEAT\|EnsemblGenome=TraesCS1D02G451300\|UniProtKB=A0A3B6A2S9	Triticum aestivum	LDO
WHEAT\|EnsemblGenome=TraesCS1B02G477500\|UniProtKB=A0A3B5Z6R9	Triticum aestivum	LDO
WHEAT\|EnsemblGenome=TraesCS1A02G443400\|UniProtKB=A0A3B5Y8V4	Triticum aestivum	LDO
NELNU\|Gene_ORFName=HUJ06_028621\|UniProtKB=A0A1U7ZK65	Nelumbo nucifera	O
NELNU\|Gene=LOC104592428\|UniProtKB=A0A1U7ZFC0	Nelumbo nucifera	LDO
HELAN\|EnsemblGenome=HanXRQr2_Chr16g0726041\|UniProtKB=A0A251RW81	Helianthus annuus	O
HELAN\|EnsemblGenome=HanXRQr2_Chr11g0520461\|UniProtKB=A0A251TFV1	Helianthus annuus	O
SOLTU\|EnsemblGenome=PGSC0003DMG400020476\|UniProtKB=M1BTY2	Solanum tuberosum	O
SOLLC\|EnsemblGenome=Solyc06g066060.3\|UniProtKB=A0A3Q7GZ89	Solanum lycopersicum	O
VITVI\|EnsemblGenome=Vitvi09g00588\|UniProtKB=D7U112	Vitis vinifera	O
RICCO\|Gene_ORFName=RCOM_0899160\|UniProtKB=B9RV50	Ricinus communis	O
JUGRE\|EnsemblGenome=Jr07_31070\|UniProtKB=A0A2I4H3C0	Juglans regia	O
JUGRE\|EnsemblGenome=Jr07_31090\|UniProtKB=A0A6P9EKN5	Juglans regia	O
JUGRE\|EnsemblGenome=Jr12_19890\|UniProtKB=A0A6P9DW83	Juglans regia	O
PRUPE\|EnsemblGenome=PRUPE_3G084000\|UniProtKB=A0A251PX97	Prunus persica	O
SPIOL\|Gene=LOC110788839\|UniProtKB=A0A9R0IHF1	Spinacia oleracea	O
SPIOL\|Gene=LOC110786772\|UniProtKB=A0A9R0IE74	Spinacia oleracea	LDO
HELAN\|Gene_ORFName=HanXRQr2_Chr16g0745461\|UniProtKB=A0A9K3DR30	Helianthus annuus	LDO
HELAN\|Gene_ORFName=HannXRQ_Chr16g0510851\|UniProtKB=A0A251RYB8	Helianthus annuus	LDO
LACSA\|Gene_ORFName=LSAT_V11C500239680\|UniProtKB=A0A9R1XBY9	Lactuca sativa	LDO
ERYGU\|Gene_ORFName=MIMGU_mgv1a006879mg\|UniProtKB=A0A022QYC3	Erythranthe guttata	LDO
CAPAN\|EnsemblGenome=T459_16435\|UniProtKB=A0A1U8GWM0	Capsicum annuum	O
TOBAC\|Gene=LOC107774696\|UniProtKB=A0A1S3YCS7	Nicotiana tabacum	O
TOBAC\|Gene=LOC107759702\|UniProtKB=A0A1S3X077	Nicotiana tabacum	O
CAPAN\|EnsemblGenome=T459_21341\|UniProtKB=A0A2G2YWF8	Capsicum annuum	LDO
SOLTU\|EnsemblGenome=PGSC0003DMG400020781\|UniProtKB=M1BV48	Solanum tuberosum	LDO
SOLLC\|EnsemblGenome=Solyc08g061000.3\|UniProtKB=A0A3Q7IIB9	Solanum lycopersicum	LDO
TOBAC\|Gene=LOC107770853\|UniProtKB=A0A1S3Y0E6	Nicotiana tabacum	LDO
TOBAC\|Gene=LOC107800056\|UniProtKB=A0A1S4APK5	Nicotiana tabacum	O
TOBAC\|Gene=LOC107813637\|UniProtKB=A0A1S4C023	Nicotiana tabacum	O
TOBAC\|Gene=LOC107767550\|UniProtKB=A0A1S3XQ54	Nicotiana tabacum	LDO
VITVI\|EnsemblGenome=Vitvi04g00474\|UniProtKB=F6H316	Vitis vinifera	LDO
EUCGR\|EnsemblGenome=EUGRSUZ_C02784\|UniProtKB=A0A059CSQ3	Eucalyptus grandis	O
EUCGR\|EnsemblGenome=EUGRSUZ_C01749\|UniProtKB=A0A059CR29	Eucalyptus grandis	LDO
BRANA\|Gene_ORFName=GSBRNA2T00024143001\|UniProtKB=A0A078GF87	Brassica napus	O
BRANA\|Gene_ORFName=DARMORV10_C02P04900.1\|UniProtKB=A0A078IMF3	Brassica napus	LDO
BRACM\|EnsemblGenome=Bra008984\|UniProtKB=M4CXN7	Brassica campestris	O
BRACM\|EnsemblGenome=Bra008983\|UniProtKB=M4CXN6	Brassica campestris	LDO
BRACM\|EnsemblGenome=Bra006091\|UniProtKB=M4CPF3	Brassica campestris	O
BRACM\|EnsemblGenome=Bra023311\|UniProtKB=M4E3F9	Brassica campestris	O
BRACM\|EnsemblGenome=Bra023312\|UniProtKB=M4E3G0	Brassica campestris	LDO
ARATH\|TAIR=locus=2147987\|UniProtKB=Q56XG6	Arabidopsis thaliana	LDO
ARATH\|TAIR=locus=2148027\|UniProtKB=Q9LFN6	Arabidopsis thaliana	LDO
GOSHI\|Gene=LOC107914165\|UniProtKB=A0A1U8KCR9	Gossypium hirsutum	O
GOSHI\|Gene=LOC107897262\|UniProtKB=A0A1U8IQI2	Gossypium hirsutum	LDO
THECC\|EnsemblGenome=TCM_027151\|UniProtKB=A0A061G7G0	Theobroma cacao	LDO
GOSHI\|Gene=LOC107940048\|UniProtKB=A0A1U8MTS4	Gossypium hirsutum	O
GOSHI\|Gene=LOC107958960\|UniProtKB=A0A1U8PGN2	Gossypium hirsutum	O
GOSHI\|Gene=LOC107893444\|UniProtKB=A0A1U8I6S7	Gossypium hirsutum	O
GOSHI\|Gene=LOC107948699\|UniProtKB=A0A1U8NIW0	Gossypium hirsutum	O
GOSHI\|Gene=LOC107901605\|UniProtKB=A0A1U8IYF0	Gossypium hirsutum	O
GOSHI\|Gene=LOC107906884\|UniProtKB=A0A1U8JLE7	Gossypium hirsutum	O
GOSHI\|Gene=LOC107888417\|UniProtKB=A0A1U8HXB4	Gossypium hirsutum	O
GOSHI\|Gene=LOC107962898\|UniProtKB=A0A1U8PUW5	Gossypium hirsutum	O
THECC\|EnsemblGenome=TCM_037013\|UniProtKB=A0A061GK01	Theobroma cacao	O
CITSI\|Gene_ORFName=CISIN_1g014314mg\|UniProtKB=A0A067F2Y2	Citrus sinensis	LDO
POPTR\|EnsemblGenome=POPTR_018G028600v3\|UniProtKB=B9IKW5	Populus trichocarpa	O
POPTR\|EnsemblGenome=POPTR_006G253100v3\|UniProtKB=B9HA46	Populus trichocarpa	LDO
MANES\|EnsemblGenome=MANES_16G121100\|UniProtKB=A0A2C9UAV4	Manihot esculenta	LDO
MANES\|EnsemblGenome=MANES_03G015600\|UniProtKB=A0A2C9W3M6	Manihot esculenta	O
SOYBN\|EnsemblGenome=GLYMA_07G031400\|UniProtKB=A0A0R0J4W7	Glycine max	LDO
SOYBN\|EnsemblGenome=GLYMA_08G211400\|UniProtKB=I1KVD4	Glycine max	LDO
MEDTR\|EnsemblGenome=MTR_4g010565\|UniProtKB=A0A072UHT1	Medicago truncatula	LDO
SOYBN\|EnsemblGenome=GLYMA_04G067600\|UniProtKB=I1JUD5	Glycine max	LDO
SOYBN\|EnsemblGenome=GLYMA_06G069200\|UniProtKB=A0A0R0JKX7	Glycine max	LDO
MEDTR\|EnsemblGenome=MTR_3g107420\|UniProtKB=G7JB35	Medicago truncatula	O
MEDTR\|EnsemblGenome=MTR_1g012420\|UniProtKB=G7I6Z7	Medicago truncatula	LDO
JUGRE\|EnsemblGenome=Jr08_01510\|UniProtKB=A0A2I4E0E7	Juglans regia	O
JUGRE\|EnsemblGenome=Jr11_29280\|UniProtKB=A0A2I4ES45	Juglans regia	LDO
CUCSA\|EnsemblGenome=Csa_2G381720\|UniProtKB=A0A0A0LSH7	Cucumis sativus	LDO
PRUPE\|EnsemblGenome=PRUPE_1G479900\|UniProtKB=M5XQ24	Prunus persica	LDO
SELML\|EnsemblGenome=SELMODRAFT_146823\|UniProtKB=D8RFZ4	Selaginella moellendorffii	LDO
SELML\|EnsemblGenome=SELMODRAFT_446187\|UniProtKB=D8SPE8	Selaginella moellendorffii	LDO
PHYPA\|EnsemblGenome=Pp3c7_5890\|UniProtKB=A0A2K1KAL0	Physcomitrella patens	LDO
MARPO\|EnsemblGenome=MARPO_0029s0012\|UniProtKB=A0A2R6X8N7	Marchantia polymorpha	LDO
KLENI\|Gene_ORFName=KFL_002100080\|UniProtKB=A0A1Y1I9V7	Klebsormidium nitens	LDO
NITMS\|EnsemblGenome=Nmar_1182\|UniProtKB=A9A5S3	Nitrosopumilus maritimus	O
SACS2\|EnsemblGenome=SSO2036\|UniProtKB=Q97WT0	Sulfolobus solfataricus	O
SACS2\|EnsemblGenome=SSO1889\|UniProtKB=Q97X64	Sulfolobus solfataricus	O