PANTHER Gene Information

Gene Symbol(s):

ULBP21

Organism:

Bos taurus

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Tree

Gene Name:

UL16 binding protein 21

Gene ID:

ENSBTAG00000047902.2

Protein ID:

Q09YM2

Persistent Id:

PTN001047485

Alternate Ids:

Q09YM2(Synonym)	ENSBTAP00000053867(Ensembl_PRO)
DP000081(EMBL)	ABI75237(EMBL-CDS)
114540274(GI)	ENSBTAG00000047902(Ensembl)
Q09YM2_BOVIN(UniProtKB-ID)	NP_001162079(refSeq)
ENSBTAT00000064751(Ensembl_TRS)	513308(GeneID)
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PANTHER Classification

PANTHER Family:	MHC CLASS I-RELATED (PTHR16675)
PANTHER Subfamily:	UL16 BINDING PROTEIN 21 (PTHR16675:SF1)
PANTHER GO-slim Molecular Function:
PANTHER GO-slim Biological Process:	immune response regulation of leukocyte mediated cytotoxicity antigen processing and presentation positive regulation of cellular process positive regulation of adaptive immune response regulation of lymphocyte mediated immunity
PANTHER GO-slim Cellular Component:	external side of plasma membrane extracellular space
PANTHER protein class:	major histocompatibility complex protein

Pathway Categories:

No pathway information available

GENE ONTOLOGY DATABASE ANNOTATIONS

Click to Expand...

GO MF Complete:	-
GO BP Complete:	immune response, positive regulation of T cell mediated cytotoxicity, antigen processing and presentation of endogenous peptide antigen via MHC class I via ER pathway, TAP-independent, antigen processing and presentation of endogenous peptide antigen via MHC class Ib
GO CC Complete:	extracellular space, external side of plasma membrane

REACTOME DATABASE ANNOTATIONS Click to Expand...

Reactome Pathways:

Adaptive Immune System, Post-translational protein modification, Metabolism of proteins, Immune System, Post-translational modification: synthesis of GPI-anchored proteins, Immunoregulatory interactions between a Lymphoid and a non-Lymphoid cell

Orthologs

ID	Organism	Type
HUMAN\|HGNC=16795\|UniProtKB=Q6H3X3	Homo sapiens	O
PANTR\|Ensembl=ENSPTRG00000029377.4\|UniProtKB=K7DPA9	Pan troglodytes	O
GORGO\|Ensembl=ENSGGOG00000016690.3\|UniProtKB=G3RKL3	Gorilla gorilla gorilla	O
HUMAN\|HGNC=16798\|UniProtKB=Q5VY80	Homo sapiens	O
HUMAN\|HGNC=14894\|UniProtKB=Q9BZM5	Homo sapiens	O
PANTR\|Ensembl=ENSPTRG00000052558.1\|UniProtKB=H2R0L5	Pan troglodytes	O
PANTR\|Ensembl=ENSPTRG00000044735.1\|UniProtKB=H2R127	Pan troglodytes	O
GORGO\|Ensembl=ENSGGOG00000007925.3\|UniProtKB=G3QY25	Gorilla gorilla gorilla	O
MACMU\|Ensembl=ENSMMUG00000053480.1\|UniProtKB=A0A5F7ZAF9	Macaca mulatta	O
MACMU\|Ensembl=ENSMMUG00000029466.3\|UniProtKB=F7FTW6	Macaca mulatta	O
MACMU\|Ensembl=ENSMMUG00000029465.3\|UniProtKB=F6R7A7	Macaca mulatta	O
MACMU\|Ensembl=ENSMMUG00000003951.4\|UniProtKB=F6ZGG1	Macaca mulatta	O
HUMAN\|HGNC=14895\|UniProtKB=Q9BZM4	Homo sapiens	O
PANTR\|Ensembl=ENSPTRG00000034286.3\|UniProtKB=H2R6W1	Pan troglodytes	O
GORGO\|Ensembl=ENSGGOG00000003891.3\|UniProtKB=G3QMS0	Gorilla gorilla gorilla	O
MACMU\|Ensembl=ENSMMUG00000047033.2\|UniProtKB=A0A1D5R722	Macaca mulatta	O
HUMAN\|HGNC=14893\|UniProtKB=Q9BZM6	Homo sapiens	O
PANTR\|Ensembl=ENSPTRG00000043696.1\|UniProtKB=H2QZY1	Pan troglodytes	O
GORGO\|Ensembl=ENSGGOG00000007707.3\|UniProtKB=G3QXJ6	Gorilla gorilla gorilla	O
GORGO\|Ensembl=ENSGGOG00000036889.1\|UniProtKB=A0A2I2ZU65	Gorilla gorilla gorilla	O
MACMU\|Ensembl=ENSMMUG00000063631.1\|UniProtKB=A0A5F8ALN9	Macaca mulatta	O
MACMU\|Ensembl=ENSMMUG00000065098.1\|UniProtKB=A0A5F7Z919	Macaca mulatta	O
MOUSE\|MGI=MGI=1925027\|UniProtKB=Q8HWA3	Mus musculus	O
PIG\|Ensembl=ENSSSCG00000026042.4\|UniProtKB=I6L5V9	Sus scrofa	O
HORSE\|Ensembl=ENSECAG00000033237.3\|UniProtKB=F7C7I4	Equus caballus	O
HORSE\|Ensembl=ENSECAG00000037408.3\|UniProtKB=A0A3Q2GS16	Equus caballus	O
HORSE\|Ensembl=ENSECAG00000059096.1\|UniProtKB=A0A9L0T4P3	Equus caballus	O
FELCA\|Ensembl=ENSFCAG00000047793.1\|UniProtKB=A0A5F5XQE8	Felis catus	O
FELCA\|Ensembl=ENSFCAG00000049506.1\|UniProtKB=A0A5F5XPF0	Felis catus	O
CANLF\|Ensembl=ENSCAFG00845019534.1\|UniProtKB=A0A8I3NZ50	Canis lupus familiaris	O
MONDO\|Ensembl=ENSMODG00000015798.3\|UniProtKB=A0A5F8G5B6	Monodelphis domestica	O
MONDO\|Ensembl=ENSMODG00000015796.3\|UniProtKB=A0A5F8GGQ6	Monodelphis domestica	O
ORNAN\|Ensembl=ENSOANG00000005224.4\|UniProtKB=F7BF28	Ornithorhynchus anatinus	O
ORNAN\|Ensembl=ENSOANG00000015040.2\|UniProtKB=A0A6I8NLZ2	Ornithorhynchus anatinus	O
ORNAN\|Ensembl=ENSOANG00000001818.4\|UniProtKB=F7GBN2	Ornithorhynchus anatinus	O
ORNAN\|Ensembl=ENSOANG00000044768.1\|UniProtKB=A0A6I8NGZ5	Ornithorhynchus anatinus	O
ORNAN\|Ensembl=ENSOANG00000001863.3\|UniProtKB=F7F763	Ornithorhynchus anatinus	O
ORNAN\|Ensembl=ENSOANG00000037190.1\|UniProtKB=A7X5N1	Ornithorhynchus anatinus	O
ORNAN\|Ensembl=ENSOANG00000003398.3\|UniProtKB=A0A6I8P8S3	Ornithorhynchus anatinus	O
CHICK\|Gene=HA1F_CHICK\|UniProtKB=P15979	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010002457.1\|UniProtKB=A0A8V0XA48	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010002333.1\|UniProtKB=A0A8V0X8Y6	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010003796.1\|UniProtKB=A0A8V0XLL9	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010002122.1\|UniProtKB=A0A8V0XEM0	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010002311.1\|UniProtKB=A0A8V0X510	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010003731.1\|UniProtKB=A0A8V0XLD0	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010003772.1\|UniProtKB=A0A8V0XHA9	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010003767.1\|UniProtKB=A0A8V0XLH9	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010003722.1\|UniProtKB=A0A8V0XLB8	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010003749.1\|UniProtKB=A0A8V0XEE7	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010003817.1\|UniProtKB=A0A089FKX6	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010002414.1\|UniProtKB=A0A8V0X9R8	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010002274.1\|UniProtKB=A0A8V0XF73	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010002423.1\|UniProtKB=A0A8V0XFJ7	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010000180.1\|UniProtKB=A0A8V0WZL7	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010003756.1\|UniProtKB=A0A8V0XJY8	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010002473.1\|UniProtKB=A0A8V0XD73	Gallus gallus	O
CHICK\|Ensembl=ENSGALG00010003110.1\|UniProtKB=A0ZXL5	Gallus gallus	O
ANOCA\|Ensembl=ENSACAG00000006471.4\|UniProtKB=H9GBH1	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000024638.3\|UniProtKB=H9GQU0	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000011968.4\|UniProtKB=H9GR22	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000042239.1\|UniProtKB=A0A803SU65	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000022634.2\|UniProtKB=L7MZR7	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000023667.2\|UniProtKB=H9GR57	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000027871.2\|UniProtKB=H9GTB3	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000035066.1\|UniProtKB=A0A803T250	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000036451.1\|UniProtKB=A0A803T973	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000022657.3\|UniProtKB=H9GUX5	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000034645.1\|UniProtKB=A0A803U071	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000003922.3\|UniProtKB=A0A803T697	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000000468.3\|UniProtKB=A0A803TQH7	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000024309.3\|UniProtKB=H9GQP3	Anolis carolinensis	O
XENLA\|Gene=mhc1b-uea.L\|UniProtKB=A0A8J0V155	Xenopus laevis	O
XENLA\|Gene=mhc100494167.L\|UniProtKB=A0A8J1KLG8	Xenopus laevis	O
XENTR\|Ensembl=ENSXETG00000039951\|UniProtKB=A0A6I8RP42	Xenopus tropicalis	O
XENLA\|Gene=LOC108699401\|UniProtKB=A0A8J1LLA4	Xenopus laevis	O
XENTR\|Ensembl=ENSXETG00000030689\|UniProtKB=K9J892	Xenopus tropicalis	O
XENLA\|Gene=LOC121402286\|UniProtKB=A0A8J1MT10	Xenopus laevis	O
XENLA\|Gene=LOC108711789\|UniProtKB=A0A1L8H432	Xenopus laevis	O
XENTR\|Xenbase=XB-GENE-5873924\|UniProtKB=A0A6I8R1D6	Xenopus tropicalis	O
XENLA\|Gene=LOC108699591\|UniProtKB=A0A8J0TE81	Xenopus laevis	O
XENTR\|Xenbase=XB-GENE-484711\|UniProtKB=A0A8J0QH19	Xenopus tropicalis	O
XENTR\|Gene=LOC100497446\|UniProtKB=A0A8J0R7Z3	Xenopus tropicalis	O
XENTR\|Gene=LOC100494930\|UniProtKB=A0A8J0R8A7	Xenopus tropicalis	O
XENTR\|Ensembl=ENSXETG00000012424\|UniProtKB=A0A6I8QD81	Xenopus tropicalis	O
XENLA\|Gene=LOC121393115\|UniProtKB=A0A8J1LP79	Xenopus laevis	O
XENTR\|Gene=LOC100496024\|UniProtKB=A0A8J1IQ11	Xenopus tropicalis	O
XENLA\|Gene=LOC108699218\|UniProtKB=A0A8J0TL10	Xenopus laevis	O
XENTR\|Gene=LOC100485167\|UniProtKB=A0A8J0QQ05	Xenopus tropicalis	O
XENLA\|Gene=LOC108699418\|UniProtKB=A0A8J0TDW6	Xenopus laevis	O
XENTR\|Gene=LOC100494130\|UniProtKB=A0A8J0R7H6	Xenopus tropicalis	O
XENLA\|Gene=LOC108700607\|UniProtKB=A0A8J1LP02	Xenopus laevis	O
XENTR\|Gene=LOC100490896\|UniProtKB=A0A8J1IQ84	Xenopus tropicalis	O
XENLA\|Gene=LOC108700369\|UniProtKB=A0A8J1LMJ5	Xenopus laevis	O
XENTR\|Ensembl=ENSXETG00000046574\|UniProtKB=A0A803JBZ8	Xenopus tropicalis	O
XENTR\|Gene=LOC100495868\|UniProtKB=A0A8J1IQ19	Xenopus tropicalis	O
XENTR\|Gene=LOC116406481\|UniProtKB=A0A8J1IQ18	Xenopus tropicalis	O
XENTR\|Gene=LOC100495225\|UniProtKB=A0A8J1IMD3	Xenopus tropicalis	O
XENTR\|Gene=LOC100494293\|UniProtKB=A0A8J0T6K7	Xenopus tropicalis	O
XENTR\|Gene=LOC116406581\|UniProtKB=A0A8J1IMD1	Xenopus tropicalis	O
XENTR\|Gene=LOC101734387\|UniProtKB=A0A8J1IMD2	Xenopus tropicalis	O
XENTR\|Gene=LOC100495542\|UniProtKB=A0A8J0QWT4	Xenopus tropicalis	O
XENTR\|Gene=LOC100494618\|UniProtKB=A0A8J1IQA6	Xenopus tropicalis	O
XENLA\|Gene=mhc1b-uba6.3.L\|UniProtKB=Q6AX04	Xenopus laevis	O
XENLA\|Gene=LOC108699400\|UniProtKB=A0A8J0TK51	Xenopus laevis	O
XENLA\|Gene=mhc1b-uba6.1\|UniProtKB=Q5HZZ1	Xenopus laevis	O
XENLA\|Gene=LOC108699590\|UniProtKB=A0A8J0TKK3	Xenopus laevis	O
XENLA\|Gene=XB5820455.L\|UniProtKB=A0A8J1LF77	Xenopus laevis	O
XENTR\|Gene=LOC100490553\|UniProtKB=A0A8J1IP45	Xenopus tropicalis	O
XENLA\|Gene=mhc1b2.L\|UniProtKB=A0A8J1LJP0	Xenopus laevis	O
XENTR\|Gene=LOC108644486\|UniProtKB=A0A8J1IPL0	Xenopus tropicalis	O
XENLA\|Gene=mr1-like.L\|UniProtKB=A0A8J0Q5V1	Xenopus laevis	O
XENLA\|Gene=LOC121397207\|UniProtKB=A0A8J1LK94	Xenopus laevis	O
XENTR\|Xenbase=XB-GENE-5961407\|UniProtKB=B2RYZ4	Xenopus tropicalis	O
XENTR\|Gene=LOC100496488\|UniProtKB=A0A8J0SW22	Xenopus tropicalis	O
XENTR\|Gene=LOC100493954\|UniProtKB=A0A8J0S5P6	Xenopus tropicalis	O
XENTR\|Gene=LOC100496176\|UniProtKB=A0A8J0SA08	Xenopus tropicalis	O
XENTR\|Xenbase=XB-GENE-5991273\|UniProtKB=A0A8J0QTS3	Xenopus tropicalis	O
XENTR\|Gene=LOC100496639\|UniProtKB=A0A8J1ISF8	Xenopus tropicalis	O
XENLA\|Gene=LOC121397201\|UniProtKB=A0A8J1LK89	Xenopus laevis	O
XENLA\|Gene=LOC121397198\|UniProtKB=A0A8J1LJN6	Xenopus laevis	O
XENLA\|Gene=LOC108699126\|UniProtKB=A0A8J1LK81	Xenopus laevis	O
XENLA\|Gene=mhc1b.L\|UniProtKB=A0A8J1LGJ4	Xenopus laevis	O
XENLA\|Gene=LOC121397208\|UniProtKB=A0A8J1LLD1	Xenopus laevis	O
XENLA\|Gene=mhc1b-uba1.1.L\|UniProtKB=A9UM13	Xenopus laevis	O
XENLA\|Gene=LOC108699125\|UniProtKB=A0A8J1LLB8	Xenopus laevis	O
XENLA\|Gene=LOC108699407\|UniProtKB=A0A8J1LIU7	Xenopus laevis	O
XENLA\|Gene=LOC108699121\|UniProtKB=A0A1L8FCD5	Xenopus laevis	O
XENLA\|Gene=LOC108699128\|UniProtKB=A0A1L8FCG0	Xenopus laevis	O
XENLA\|Gene=LOC108699124\|UniProtKB=A0A8J1LJ07	Xenopus laevis	O
XENLA\|Gene=LOC108699123\|UniProtKB=A0A8J0TCY5	Xenopus laevis	O
XENLA\|Gene=LOC121397200\|UniProtKB=A0A8J1LJ12	Xenopus laevis	O
XENLA\|Gene=LOC108705562\|UniProtKB=A0A8J1LIV7	Xenopus laevis	O
XENTR\|Gene=LOC100495703\|UniProtKB=A0A8J0QWH3	Xenopus tropicalis	O
XENTR\|Gene=LOC113455423\|UniProtKB=A0A8J0QHA2	Xenopus tropicalis	O
XENLA\|Gene=LOC108698552\|UniProtKB=A0A8J1LHH5	Xenopus laevis	O
XENTR\|Xenbase=XB-GENE-485307\|UniProtKB=Q6XXY3	Xenopus tropicalis	O
DANRE\|ZFIN=ZDB-GENE-140820-4\|UniProtKB=A0A0R4IZK4	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-140820-11\|UniProtKB=X1WF83	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-140820-8\|UniProtKB=F1RAH2	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-140820-5\|UniProtKB=F1RAH3	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-140820-13\|UniProtKB=A0A0G2KIH5	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-161101-1\|UniProtKB=X1WF74	Danio rerio	O
DANRE\|Gene=mhc1lba\|UniProtKB=A0A8M3AVG2	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-050417-464\|UniProtKB=Q566P2	Danio rerio	O
DANRE\|Gene=mhc1lea\|UniProtKB=A0A8M9PVR9	Danio rerio	O
DANRE\|Gene=si=dkey-225f5.5\|UniProtKB=A0A8M9PDN6	Danio rerio	O
DANRE\|Gene=mhc1lfa\|UniProtKB=A0A8M1P987	Danio rerio	O
DANRE\|Gene=LOC103909824\|UniProtKB=A0A8M9PNJ1	Danio rerio	O
DANRE\|Gene=LOC108181585\|UniProtKB=A0A8M9PZC0	Danio rerio	O
LEPOC\|Ensembl=ENSLOCG00000000335.1\|UniProtKB=W5LWA2	lepisosteus oculatus	O
LEPOC\|Ensembl=ENSLOCG00000000528.1\|UniProtKB=W5LWS5	lepisosteus oculatus	O
LEPOC\|Ensembl=ENSLOCG00000000416.1\|UniProtKB=W5LWF3	lepisosteus oculatus	O
ORYLA\|Ensembl=ENSORLG00000024564.1\|UniProtKB=H2L676	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000001056.2\|UniProtKB=H2L659	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000001044.2\|UniProtKB=H2N1H0	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000024388.1\|UniProtKB=A0A3B3HSY5	Oryzias latipes	O
DANRE\|ZFIN=ZDB-GENE-070424-4\|UniProtKB=A4FUM4	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-070705-176\|UniProtKB=A8E7F2	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-040426-2149\|UniProtKB=A8E7F4	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-130206-9\|UniProtKB=U6BNE6	Danio rerio	O
DANRE\|Gene=mhc1zja\|UniProtKB=A0A8M2BIV1	Danio rerio	O
DANRE\|Gene=LOC571647\|UniProtKB=A7MBX5	Danio rerio	O
DANRE\|Gene=LOC563036\|UniProtKB=A0A8M1QUM8	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-130206-7\|UniProtKB=U6BMG4	Danio rerio	O
ORYLA\|Ensembl=ENSORLG00000006575.2\|UniProtKB=A0A3B3HEA8	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000006798.2\|UniProtKB=H2LR16	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000018729.2\|UniProtKB=H2MWX4	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000000983.2\|UniProtKB=H2L5W4	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000017153.2\|UniProtKB=F5HRI2	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000020349.2\|UniProtKB=H2N1C6	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000029335.1\|UniProtKB=H2MAI6	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000000970.2\|UniProtKB=H2L5U7	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000019579.2\|UniProtKB=A0A3B3IH85	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000012414.2\|UniProtKB=H2MAI9	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000006435.2\|UniProtKB=A0A3B3H8R3	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000006772.2\|UniProtKB=H2LR18	Oryzias latipes	O
DANRE\|ZFIN=ZDB-GENE-140820-1\|UniProtKB=E7FAH6	Danio rerio	O
DANRE\|Gene=mhc1ula\|UniProtKB=A0A0S1RUH3	Danio rerio	O
DANRE\|Gene=mhc1uka\|UniProtKB=A0A8M2B625	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-990415-145\|UniProtKB=Q31365	Danio rerio	O
LEPOC\|Ensembl=ENSLOCG00000008666.1\|UniProtKB=W5MQ86	lepisosteus oculatus	O
LEPOC\|Ensembl=ENSLOCG00000000215.1\|UniProtKB=W5LVT1	lepisosteus oculatus	O
LEPOC\|Ensembl=ENSLOCG00000000807.1\|UniProtKB=W5LXP7	lepisosteus oculatus	O
LEPOC\|Ensembl=ENSLOCG00000007238.1\|UniProtKB=W5MK52	lepisosteus oculatus	O