PANTHER Gene Information

Gene Symbol(s):

MRS4

Organism:

Candida albicans

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Gene Name:

Mitochondrial thiamine pyrophosphate carrier 1

Gene ID:

CAL0000193287

Protein ID:

Q5A2T7

Persistent Id:

PTN001183486

Alternate Ids:

C2_08070C_A-T-p1(Ensembl_PRO)	Q5A2T7_CANAL(UniProtKB-ID)
C2_08070C_A(Ensembl)	CP017624(EMBL)
1078719256(GI)	AOW27801(EMBL-CDS)
XM_710926(refSeq)	C2_08070C_A-T(Ensembl_TRS)
3642306(GeneID)	Q5A2T7(Synonym)
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PANTHER Classification

PANTHER Family:	MITOFERRIN-1-RELATED (PTHR45758)
PANTHER Subfamily:	MITOFERRIN-1 (PTHR45758:SF4)
PANTHER GO-slim Molecular Function:	metal ion transmembrane transporter activity
PANTHER GO-slim Biological Process:	mitochondrial transmembrane transport iron ion transmembrane transport
PANTHER GO-slim Cellular Component:	mitochondrial membrane
PANTHER protein class:

Pathway Categories:

No pathway information available

GENE ONTOLOGY DATABASE ANNOTATIONS

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GO MF Complete:	iron ion transmembrane transporter activity, ferrous iron transmembrane transporter activity
GO BP Complete:	intracellular iron ion homeostasis, transmembrane transport, iron import into the mitochondrion
GO CC Complete:	mitochondrial membrane, mitochondrion, mitochondrial inner membrane, plasma membrane

REACTOME DATABASE ANNOTATIONS Click to Expand...

Reactome Pathways:

Orthologs

ID	Organism	Type
HUMAN\|HGNC=29786\|UniProtKB=Q9NYZ2	Homo sapiens	LDO
PANTR\|Ensembl=ENSPTRG00000020079.5\|UniProtKB=A0A2I3T209	Pan troglodytes	LDO
GORGO\|Ensembl=ENSGGOG00000005025.3\|UniProtKB=A0A2I2ZXB2	Gorilla gorilla gorilla	LDO
MACMU\|Ensembl=ENSMMUG00000054404.1\|UniProtKB=A0A5F8AQR0	Macaca mulatta	LDO
MOUSE\|MGI=MGI=1914962\|UniProtKB=Q920G8	Mus musculus	LDO
RAT\|RGD=1359361\|UniProtKB=Q66H23	Rattus norvegicus	LDO
BOVIN\|Ensembl=ENSBTAG00000019099.5\|UniProtKB=E1B7M3	Bos taurus	LDO
PIG\|Ensembl=ENSSSCG00000033699.1\|UniProtKB=A0A8W4FMP0	Sus scrofa	O
PIG\|Ensembl=ENSSSCG00000033037.2\|UniProtKB=A0A287ABD4	Sus scrofa	LDO
HORSE\|Ensembl=ENSECAG00000013319.4\|UniProtKB=A0A3Q2GS60	Equus caballus	LDO
FELCA\|Ensembl=ENSFCAG00000005418.6\|UniProtKB=M3W411	Felis catus	LDO
CANLF\|Ensembl=ENSCAFG00845016130.1\|UniProtKB=A0A8I3NU89	Canis lupus familiaris	LDO
ORNAN\|Ensembl=ENSOANG00000008028.2\|UniProtKB=F7ENY9	Ornithorhynchus anatinus	LDO
CHICK\|Ensembl=ENSGALG00010020244.1\|UniProtKB=F1NS09	Gallus gallus	LDO
ANOCA\|Ensembl=ENSACAG00000012023.3\|UniProtKB=H9GIE5	Anolis carolinensis	LDO
XENLA\|Gene=slc25a37.S\|UniProtKB=A0A1L8GRS8	Xenopus laevis	LDO
XENLA\|Gene=slc25a37.L\|UniProtKB=A0A1L8H1T2	Xenopus laevis	LDO
XENTR\|Xenbase=XB-GENE-999414\|UniProtKB=F7ETQ6	Xenopus tropicalis	LDO
ORYLA\|Ensembl=ENSORLG00000010917.2\|UniProtKB=H2M5G3	Oryzias latipes	LDO
DANRE\|ZFIN=ZDB-GENE-031118-202\|UniProtKB=Q287T7	Danio rerio	LDO
LEPOC\|Ensembl=ENSLOCG00000015013.1\|UniProtKB=W5NCX3	lepisosteus oculatus	LDO
HELRO\|EnsemblGenome=HelroG185413\|UniProtKB=T1FMS4	helobdella robusta	LDO
HUMAN\|HGNC=23472\|UniProtKB=Q96A46	Homo sapiens	O
PANTR\|Ensembl=ENSPTRG00000002831.4\|UniProtKB=H2Q2E6	Pan troglodytes	O
GORGO\|Ensembl=ENSGGOG00000000150.3\|UniProtKB=G3QD45	Gorilla gorilla gorilla	O
MACMU\|Ensembl=ENSMMUG00000023760.4\|UniProtKB=F7EJV4	Macaca mulatta	O
MOUSE\|MGI=MGI=2180509\|UniProtKB=Q8R0Z5	Mus musculus	O
RAT\|RGD=1584155\|UniProtKB=G3V865	Rattus norvegicus	O
BOVIN\|Ensembl=ENSBTAG00000012107.5\|UniProtKB=E1BJE6	Bos taurus	O
PIG\|Ensembl=ENSSSCG00000037536.2\|UniProtKB=A0A286ZJV0	Sus scrofa	O
HORSE\|Ensembl=ENSECAG00000009356.4\|UniProtKB=F6WZG2	Equus caballus	O
FELCA\|Ensembl=ENSFCAG00000045981.2\|UniProtKB=A0A337S4B3	Felis catus	O
CANLF\|Ensembl=ENSCAFG00845030792.1\|UniProtKB=A0A8I3Q7U8	Canis lupus familiaris	O
MONDO\|Ensembl=ENSMODG00000003519.4\|UniProtKB=F7GBV9	Monodelphis domestica	O
ORNAN\|Ensembl=ENSOANG00000006335.3\|UniProtKB=F6R5Y3	Ornithorhynchus anatinus	O
CHICK\|Ensembl=ENSGALG00010022250.1\|UniProtKB=A0A8V0ZV01	Gallus gallus	O
ANOCA\|Ensembl=ENSACAG00000003687.3\|UniProtKB=G1KC52	Anolis carolinensis	O
XENLA\|Gene=LOC121395366\|UniProtKB=A0A8J1L7V7	Xenopus laevis	O
XENLA\|Gene=LOC108697466\|UniProtKB=A0A8J0TE07	Xenopus laevis	O
XENTR\|Xenbase=XB-GENE-967691\|UniProtKB=A0JM88	Xenopus tropicalis	O
ORYLA\|Ensembl=ENSORLG00000023276.1\|UniProtKB=A0A3B3IMW3	Oryzias latipes	O
ORYLA\|Ensembl=ENSORLG00000007710.2\|UniProtKB=A0A3B3I205	Oryzias latipes	O
DANRE\|ZFIN=ZDB-GENE-040426-2125\|UniProtKB=Q7T292	Danio rerio	O
LEPOC\|Ensembl=ENSLOCG00000009024.1\|UniProtKB=W5MRI5	lepisosteus oculatus	O
CIOIN\|Ensembl=ENSCING00000008166.3\|UniProtKB=F6UQ99	Ciona intestinalis	O
BRAFL\|Gene_ORFName=BRAFLDRAFT_124272\|UniProtKB=C3XZT1	Branchiostoma floridae	O
STRPU\|EnsemblGenome=LOC584279\|UniProtKB=A0A7M7RCS0	Strongylocentrotus purpuratus	O
CAEEL\|WormBase=WBGene00012204\|UniProtKB=Q23125	Caenorhabditis elegans	O
CAEBR\|WormBase=WBGene00025852\|UniProtKB=Q620A6	Caenorhabditis briggsae	O
PRIPA\|EnsemblGenome=WBGene00111892\|UniProtKB=A0A2A6CY07	Pristionchus pacificus	O
DROME\|FlyBase=FBgn0039561\|UniProtKB=Q9VAY3	Drosophila melanogaster	O
ANOGA\|EnsemblGenome=AGAP006030\|UniProtKB=A0A1S4GRT4	Anopheles gambiae	O
TRICA\|EnsemblGenome=TC012585\|UniProtKB=D6X3C2	Tribolium castaneum	O
DAPPU\|EnsemblGenome=DAPPUDRAFT_197963\|UniProtKB=E9GPS8	Daphnia pulex	O
DAPPU\|EnsemblGenome=DAPPUDRAFT_197988\|UniProtKB=E9GPY2	Daphnia pulex	O
IXOSC\|EnsemblGenome=ISCW018156\|UniProtKB=B7PJK7	Ixodes scapularis	O
HELRO\|EnsemblGenome=HelroG187155\|UniProtKB=T1FP73	helobdella robusta	O
NEMVE\|EnsemblGenome=NEMVEDRAFT_v1g238240\|UniProtKB=A7RH70	Nematostella vectensis	LDO
TRIAD\|EnsemblGenome=TriadG32519\|UniProtKB=B3SB63	Trichoplax adhaerens	LDO
MONBE\|EnsemblGenome=MONBRDRAFT_37999\|UniProtKB=A9V539	Monosiga brevicollis	LDO
YEAST\|SGD=S000003669\|UniProtKB=P10566	Saccharomyces cerevisiae	O
YEAST\|SGD=S000001760\|UniProtKB=P23500	Saccharomyces cerevisiae	LDO
EREGS\|EnsemblGenome=AGOS_ACR260W\|UniProtKB=Q75BL1	Eremothecium gossypii	LDO
YARLI\|EnsemblGenome=YALI0_B21604g\|UniProtKB=Q6CDS3	Yarrowia lipolytica	LDO
EMENI\|EnsemblGenome=ANIA_03690\|UniProtKB=C8V7L7	Emericella nidulans	LDO
ASPFU\|EnsemblGenome=AFUA_6G12550\|UniProtKB=Q4WLR9	Neosartorya fumigata	LDO
NEUCR\|EnsemblGenome=NCU02423\|UniProtKB=Q7S4D6	Neurospora crassa	LDO
SCLS1\|Gene_ORFName=SS1G_07954\|UniProtKB=A7ERK0	Sclerotinia sclerotiorum	LDO
PHANO\|EnsemblGenome=SNOG_14516\|UniProtKB=Q0U100	Phaeosphaeria nodorum	LDO
SCHPO\|PomBase=SPAC8C9.12c\|UniProtKB=O14281	Schizosaccharomyces pombe	LDO
SCHJY\|EnsemblGenome=SJAG_02473\|UniProtKB=B6K2K4	Schizosaccharomyces japonicus	LDO
CRYNJ\|EnsemblGenome=CNE00440\|UniProtKB=Q5KHD3	Cryptococcus neoformans	LDO
USTMA\|EnsemblGenome=UMAG_01059\|UniProtKB=A0A0D1E591	Ustilago maydis	LDO
PUCGT\|EnsemblGenome=PGTG_11026\|UniProtKB=E3KN61	Puccinia graminis	LDO
BATDJ\|Gene_ORFName=BATDEDRAFT_31115\|UniProtKB=F4NUQ7	Batrachochytrium dendrobatidis	LDO
DICDI\|dictyBase=DDB_G0269470\|UniProtKB=Q55DY8	Dictyostelium discoideum	LDO
DICPU\|EnsemblGenome=DICPUDRAFT_76707\|UniProtKB=F0ZEE1	Dictyostelium purpureum	LDO
LEIMA\|EnsemblGenome=LMJF_35_3990\|UniProtKB=E9AFM4	Leishmania major	O
TRYB2\|Gene_ORFName=Tb09.211.2370\|UniProtKB=Q38DS0	Trypanosoma brucei	O
PLAF7\|EnsemblGenome=PF3D7_0108400.2\|UniProtKB=A0A143ZV51	Plasmodium falciparum	O
PLAF7\|EnsemblGenome=PF3D7_0108400.1\|UniProtKB=B9ZSI7	Plasmodium falciparum	O
PARTE\|EnsemblGenome=GSPATT00037065001\|UniProtKB=A0CBW5	Paramecium tetraurelia	O
LEIMA\|EnsemblGenome=LMJF_29_2780\|UniProtKB=E9AEE0	Leishmania major	LDO
TRYB2\|Gene_ORFName=Tb927.3.2980\|UniProtKB=Q57ZL4	Trypanosoma brucei	LDO
PLAF7\|EnsemblGenome=PF3D7_0905200\|UniProtKB=Q8I397	Plasmodium falciparum	LDO
PARTE\|EnsemblGenome=GSPATT00011091001\|UniProtKB=A0CXE2	Paramecium tetraurelia	O
PARTE\|EnsemblGenome=GSPATT00011589001\|UniProtKB=A0CYX1	Paramecium tetraurelia	LDO
THAPS\|EnsemblGenome=THAPSDRAFT_16184\|UniProtKB=B8CGM3	Thalassiosira pseudonana	LDO
PHYRM\|Gene=H3GPZ2_PHYRM\|UniProtKB=H3GPZ2	Phytophthora ramorum	LDO
CHLRE\|EnsemblGenome=CHLRE_16g695950v5\|UniProtKB=A0A2K3CSA9	Chlamydomonas reinhardtii	LDO
AMBTC\|EnsemblGenome=AMTR_s00153p00093350\|UniProtKB=W1PMK3	Amborella trichopoda	O
AMBTC\|EnsemblGenome=AMTR_s00067p00111230\|UniProtKB=U5D8L5	Amborella trichopoda	O
AMBTC\|EnsemblGenome=AMTR_s00036p00123720\|UniProtKB=U5D1Q5	Amborella trichopoda	LDO
ZOSMR\|Gene_ORFName=ZOSMA_92G00160\|UniProtKB=A0A0K9NJ68	Zostera marina	O
ZOSMR\|Gene_ORFName=ZOSMA_92G00140\|UniProtKB=A0A0K9NKR7	Zostera marina	O
MUSAM\|EnsemblGenome=Ma06_g24470\|UniProtKB=A0A804JJX7	Musa acuminata subsp. malaccensis	O
MUSAM\|EnsemblGenome=Ma03_g19030\|UniProtKB=A0A804IDS2	Musa acuminata subsp. malaccensis	O
MUSAM\|EnsemblGenome=Ma10_g13250\|UniProtKB=A0A804KVQ0	Musa acuminata subsp. malaccensis	O
SETIT\|EnsemblGenome=SETIT_036537mg\|UniProtKB=K4ACD0	Setaria italica	O
SORBI\|EnsemblGenome=SORBI_3001G403700\|UniProtKB=A0A1Z5S9Z6	Sorghum bicolor	O
MAIZE\|EnsemblGenome=Zm00001eb396750\|UniProtKB=B7ZYG8	Zea mays	O
MAIZE\|EnsemblGenome=Zm00001eb013730\|UniProtKB=B4FYW8	Zea mays	O
ORYSJ\|EnsemblGenome=Os03g0296800\|UniProtKB=Q10MS8	Oryza sativa	O
BRADI\|EnsemblGenome=BRADI_1g65170v3\|UniProtKB=A0A0Q3NWZ0	Brachypodium distachyon	O
HORVV\|EnsemblGenome=HORVU.MOREX.r3.4HG0383910\|UniProtKB=A0A8I6XKN4	Hordeum vulgare subsp. vulgare	O
WHEAT\|EnsemblGenome=TraesCS4D02G191700\|UniProtKB=A0A3B6JKY8	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS4B02G190300\|UniProtKB=A0A3B6ITF6	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS4A02G113800\|UniProtKB=A0A3B6HR23	Triticum aestivum	O
NELNU\|Gene=LOC104597279\|UniProtKB=A0A1U8A6M5	Nelumbo nucifera	O
NELNU\|Gene=LOC104602520\|UniProtKB=A0A1U8ACF2	Nelumbo nucifera	O
SPIOL\|Gene=LOC110786773\|UniProtKB=A0A9R0ID06	Spinacia oleracea	O
SPIOL\|Gene=LOC110777287\|UniProtKB=A0A9R0JKE9	Spinacia oleracea	O
HELAN\|Gene_ORFName=HanXRQr2_Chr01g0017281\|UniProtKB=A0A9K3JUK5	Helianthus annuus	O
LACSA\|Gene_ORFName=LSAT_V11C800434530\|UniProtKB=A0A9R1ULP8	Lactuca sativa	O
LACSA\|Gene_ORFName=LSAT_V11C500292350\|UniProtKB=A0A9R1VF19	Lactuca sativa	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a009856mg\|UniProtKB=A0A022S283	Erythranthe guttata	O
CAPAN\|EnsemblGenome=T459_33225\|UniProtKB=A0A2G2XZM7	Capsicum annuum	O
SOLTU\|EnsemblGenome=PGSC0003DMG400030453\|UniProtKB=M1CZQ7	Solanum tuberosum	O
SOLLC\|EnsemblGenome=Solyc06g082290.3\|UniProtKB=A0A3Q7H1Y3	Solanum lycopersicum	O
TOBAC\|Gene=LOC107782515\|UniProtKB=A0A1S3Z382	Nicotiana tabacum	O
TOBAC\|Gene=LOC107823760\|UniProtKB=A0A1S4CXV1	Nicotiana tabacum	O
TOBAC\|Gene=LOC107793347\|UniProtKB=A0A1S4A3F1	Nicotiana tabacum	O
VITVI\|EnsemblGenome=Vitvi06g00720\|UniProtKB=D7SJQ4	Vitis vinifera	O
EUCGR\|EnsemblGenome=EUGRSUZ_B03998\|UniProtKB=A0A059DAX6	Eucalyptus grandis	O
EUCGR\|EnsemblGenome=EUGRSUZ_J01862\|UniProtKB=A0A059AFH5	Eucalyptus grandis	O
BRANA\|Gene_ORFName=DARMORV10_A05P15260.1\|UniProtKB=A0A078FBH2	Brassica napus	O
BRACM\|EnsemblGenome=Bra018332\|UniProtKB=A0A397ZGP8	Brassica campestris	O
BRANA\|Gene_ORFName=DARMORV10_A04P22940.1\|UniProtKB=A0A078GYM3	Brassica napus	O
BRACM\|EnsemblGenome=Bra021617\|UniProtKB=M4DYM1	Brassica campestris	O
ARATH\|TAIR=locus=2060811\|UniProtKB=O64731	Arabidopsis thaliana	O
BRANA\|Gene_ORFName=DARMORV10_C05P05120.1\|UniProtKB=A0A078FB11	Brassica napus	O
BRACM\|EnsemblGenome=Bra015545\|UniProtKB=M4DGC2	Brassica campestris	O
ARATH\|TAIR=locus=2007387\|UniProtKB=Q8L6Z2	Arabidopsis thaliana	O
GOSHI\|Gene=LOC107963859\|UniProtKB=A0A1U8PZS2	Gossypium hirsutum	O
GOSHI\|Gene=LOC107958531\|UniProtKB=A0A1U8PHQ6	Gossypium hirsutum	O
GOSHI\|Gene=LOC107936042\|UniProtKB=A0A1U8MF16	Gossypium hirsutum	O
GOSHI\|Gene=LOC107906296\|UniProtKB=A0A1U8JKB4	Gossypium hirsutum	O
THECC\|EnsemblGenome=TCM_042220\|UniProtKB=A0A061GYQ1	Theobroma cacao	O
CITSI\|Gene_ORFName=CISIN_1g020120mg\|UniProtKB=A0A067FLK6	Citrus sinensis	O
POPTR\|EnsemblGenome=POPTR_001G282400v3\|UniProtKB=A0A2K2C518	Populus trichocarpa	O
POPTR\|EnsemblGenome=POPTR_009G077600v3\|UniProtKB=A0A2K1Z4H6	Populus trichocarpa	O
MANES\|EnsemblGenome=MANES_02G216500\|UniProtKB=A0A2C9WIE9	Manihot esculenta	O
MANES\|EnsemblGenome=MANES_18G142900\|UniProtKB=A0A2C9U514	Manihot esculenta	O
RICCO\|Gene_ORFName=RCOM_1402800\|UniProtKB=B9RRI4	Ricinus communis	O
SOYBN\|EnsemblGenome=GLYMA_02G231200\|UniProtKB=I1JHH6	Glycine max	O
SOYBN\|EnsemblGenome=GLYMA_14G198800\|UniProtKB=I1MBG1	Glycine max	O
MEDTR\|EnsemblGenome=MTR_5g073040\|UniProtKB=G7JZN0	Medicago truncatula	O
JUGRE\|EnsemblGenome=Jr14_08280\|UniProtKB=A0A2I4F9W7	Juglans regia	O
JUGRE\|EnsemblGenome=Jr14_08290\|UniProtKB=A0A2I4F9V8	Juglans regia	O
JUGRE\|Gene=LOC108989879\|UniProtKB=A0A2I4EIG3	Juglans regia	O
CUCSA\|EnsemblGenome=Csa_6G181520\|UniProtKB=A0A0A0KF81	Cucumis sativus	O
PRUPE\|EnsemblGenome=PRUPE_6G049200\|UniProtKB=M5W5R0	Prunus persica	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a026465mg\|UniProtKB=A0A022QNY2	Erythranthe guttata	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a023029mg\|UniProtKB=A0A022QHD9	Erythranthe guttata	O
VITVI\|EnsemblGenome=Vitvi08g00766\|UniProtKB=F6GXT7	Vitis vinifera	O
EUCGR\|EnsemblGenome=EUGRSUZ_K02960\|UniProtKB=A0A059A6J8	Eucalyptus grandis	O
GOSHI\|Gene=LOC107937072\|UniProtKB=A0A1U8MEW8	Gossypium hirsutum	O
THECC\|EnsemblGenome=TCM_021607\|UniProtKB=A0A061ER04	Theobroma cacao	O
CITSI\|Gene_ORFName=CISIN_1g024275mg\|UniProtKB=A0A067FIJ1	Citrus sinensis	O
POPTR\|EnsemblGenome=POPTR_016G125200v3\|UniProtKB=A0A2K1XEV3	Populus trichocarpa	O
MANES\|EnsemblGenome=MANES_08G096000\|UniProtKB=A0A2C9VEW3	Manihot esculenta	O
RICCO\|Gene_ORFName=RCOM_0847690\|UniProtKB=B9SL02	Ricinus communis	O
JUGRE\|Gene=LOC108982304\|UniProtKB=A0A2I4DPV6	Juglans regia	O
CUCSA\|EnsemblGenome=Csa_4G312860\|UniProtKB=A0A0A0L006	Cucumis sativus	O
PRUPE\|EnsemblGenome=PRUPE_6G283800\|UniProtKB=M5W051	Prunus persica	O
MUSAM\|EnsemblGenome=Ma02_g08300\|UniProtKB=A0A804I0K3	Musa acuminata subsp. malaccensis	LDO
SELML\|EnsemblGenome=SELMODRAFT_230023\|UniProtKB=D8QRH6	Selaginella moellendorffii	O
SELML\|EnsemblGenome=SELMODRAFT_232372\|UniProtKB=D8RTX5	Selaginella moellendorffii	O
SELML\|EnsemblGenome=SELMODRAFT_166450\|UniProtKB=D8QYS3	Selaginella moellendorffii	LDO
SELML\|EnsemblGenome=SELMODRAFT_95032\|UniProtKB=D8RJD2	Selaginella moellendorffii	O
SELML\|EnsemblGenome=SELMODRAFT_76024\|UniProtKB=D8QRH2	Selaginella moellendorffii	O
SELML\|EnsemblGenome=SELMODRAFT_8423\|UniProtKB=D8TGN0	Selaginella moellendorffii	O
SELML\|EnsemblGenome=SELMODRAFT_169433\|UniProtKB=D8RAD4	Selaginella moellendorffii	LDO
PHYPA\|EnsemblGenome=Pp3c12_8670\|UniProtKB=A0A2K1JPY9	Physcomitrella patens	O
PHYPA\|EnsemblGenome=Pp3c15_17240\|UniProtKB=A0A2K1JDG7	Physcomitrella patens	LDO
PHYPA\|EnsemblGenome=Pp3c9_5610\|UniProtKB=A0A7I4EQZ7	Physcomitrella patens	LDO
MARPO\|EnsemblGenome=MARPO_0047s0096\|UniProtKB=A0A2R6WZ34	Marchantia polymorpha	LDO
KLENI\|Gene_ORFName=KFL_001110110\|UniProtKB=A0A1Y1HUY3	Klebsormidium nitens	LDO