PANTHER Gene Information

Gene Symbol(s):

HannXRQ_Chr04g0113951

Organism:

Helianthus annuus

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Gene Name:

Putative bax inhibitor 1-related protein

Gene ID:

HanXRQr2_Chr04g0172021

Protein ID:

A0A251V073

Persistent Id:

PTN004627210

Alternate Ids:

KAF5810642(EMBL-CDS)	A0A251V073(Synonym)
HanXRQr2_Chr04g0172021(Ensembl)	MNCJ02000319(EMBL)
A0A251V073_HELAN(UniProtKB-ID)
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PANTHER Classification

PANTHER Family:	BAX INHIBITOR-RELATED (PTHR23291)
PANTHER Subfamily:	BAX INHIBITOR 1-RELATED PROTEIN-RELATED (PTHR23291:SF95)
PANTHER GO-slim Molecular Function:
PANTHER GO-slim Biological Process:
PANTHER GO-slim Cellular Component:	membrane
PANTHER protein class:	ion channel

Pathway Categories:

No pathway information available

GENE ONTOLOGY DATABASE ANNOTATIONS

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GO MF Complete:	-
GO BP Complete:	-
GO CC Complete:	membrane

REACTOME DATABASE ANNOTATIONS Click to Expand...

Reactome Pathways:

Orthologs

ID	Organism	Type
HUMAN\|HGNC=24257\|UniProtKB=Q9HC24	Homo sapiens	O
PANTR\|Ensembl=ENSPTRG00000005186.7\|UniProtKB=K6ZBP7	Pan troglodytes	O
GORGO\|Ensembl=ENSGGOG00000000985.3\|UniProtKB=A0A2I2YGH1	Gorilla gorilla gorilla	O
MACMU\|Ensembl=ENSMMUG00000016621.4\|UniProtKB=F6TQD4	Macaca mulatta	O
MOUSE\|MGI=MGI=1915462\|UniProtKB=Q9DA39	Mus musculus	O
RAT\|RGD=735173\|UniProtKB=Q6P9T9	Rattus norvegicus	O
BOVIN\|Ensembl=ENSBTAG00000026993.5\|UniProtKB=Q58DU1	Bos taurus	O
PIG\|Ensembl=ENSSSCG00000000473.5\|UniProtKB=A0A5G2QQH0	Sus scrofa	O
HORSE\|Ensembl=ENSECAG00000006721.4\|UniProtKB=F6UH03	Equus caballus	O
FELCA\|Ensembl=ENSFCAG00000012627.6\|UniProtKB=M3WHA1	Felis catus	O
CANLF\|Ensembl=ENSCAFG00845012562.1\|UniProtKB=A0A8I3RZH8	Canis lupus familiaris	O
MONDO\|Ensembl=ENSMODG00000009107.3\|UniProtKB=F7FCA6	Monodelphis domestica	O
ORNAN\|Ensembl=ENSOANG00000005922.3\|UniProtKB=F6QM35	Ornithorhynchus anatinus	O
CHICK\|Ensembl=ENSGALG00010012078.1\|UniProtKB=E1C1Z9	Gallus gallus	O
ANOCA\|Ensembl=ENSACAG00000003527.4\|UniProtKB=H9G7N2	Anolis carolinensis	O
XENLA\|Gene=tmbim4.L\|UniProtKB=Q5EAW5	Xenopus laevis	O
XENTR\|Xenbase=XB-GENE-947555\|UniProtKB=Q6DJB5	Xenopus tropicalis	O
DANRE\|ZFIN=ZDB-GENE-040426-2152\|UniProtKB=Q5TZH2	Danio rerio	O
LEPOC\|Ensembl=ENSLOCG00000016160.1\|UniProtKB=W5NH07	lepisosteus oculatus	O
CIOIN\|Ensembl=ENSCING00000012821.2\|UniProtKB=F6W3K8	Ciona intestinalis	O
BRAFL\|Gene_ORFName=BRAFLDRAFT_115358\|UniProtKB=C3YAB7	Branchiostoma floridae	O
STRPU\|EnsemblGenome=LOC753840\|UniProtKB=A0A7M7MZ94	Strongylocentrotus purpuratus	O
CAEEL\|WormBase=WBGene00006479\|UniProtKB=Q11080	Caenorhabditis elegans	O
CAEBR\|Gene_ORFName=CBG00173\|UniProtKB=B0K057	Caenorhabditis briggsae	O
PRIPA\|EnsemblGenome=WBGene00091150\|UniProtKB=A0A8R1Y5E4	Pristionchus pacificus	O
PRIPA\|EnsemblGenome=WBGene00114482\|UniProtKB=A0A2A6CFS1	Pristionchus pacificus	O
DROME\|FlyBase=FBgn0085459\|UniProtKB=A8DY63	Drosophila melanogaster	O
TRICA\|EnsemblGenome=TC013429\|UniProtKB=D6WLJ9	Tribolium castaneum	O
IXOSC\|EnsemblGenome=ISCW021971\|UniProtKB=B7QFX9	Ixodes scapularis	O
NEMVE\|EnsemblGenome=NEMVEDRAFT_v1g159748\|UniProtKB=A7RKF0	Nematostella vectensis	O
TRIAD\|EnsemblGenome=TriadG49581\|UniProtKB=B3RJC2	Trichoplax adhaerens	O
MONBE\|EnsemblGenome=MONBRDRAFT_25917\|UniProtKB=A9V0U8	Monosiga brevicollis	O
YEAST\|SGD=S000005249\|UniProtKB=P48558	Saccharomyces cerevisiae	O
CANAL\|CGD=CAL0000175245\|UniProtKB=A0A1D8PT90	Candida albicans	O
YARLI\|EnsemblGenome=YALI0_B05280g\|UniProtKB=Q6CFN9	Yarrowia lipolytica	O
EMENI\|EnsemblGenome=ANIA_01133\|UniProtKB=G5EB77	Emericella nidulans	O
ASPFU\|EnsemblGenome=AFUA_5G09310\|UniProtKB=Q4WUQ6	Neosartorya fumigata	O
NEUCR\|EnsemblGenome=NCU02463\|UniProtKB=Q7SG99	Neurospora crassa	O
SCLS1\|EnsemblGenome=SS1G_03316\|UniProtKB=A7EDC6	Sclerotinia sclerotiorum	O
PHANO\|EnsemblGenome=SNOG_02830\|UniProtKB=Q0UZI4	Phaeosphaeria nodorum	O
SCHPO\|PomBase=SPCC576.04\|UniProtKB=O74888	Schizosaccharomyces pombe	O
SCHJY\|EnsemblGenome=SJAG_04471\|UniProtKB=B6K6X1	Schizosaccharomyces japonicus	O
CRYNJ\|EnsemblGenome=CNG04050\|UniProtKB=Q5KDG7	Cryptococcus neoformans	O
USTMA\|EnsemblGenome=UMAG_01063\|UniProtKB=A0A0D1CXS3	Ustilago maydis	O
PUCGT\|EnsemblGenome=PGTG_09266\|UniProtKB=E3KG54	Puccinia graminis	O
DICDI\|dictyBase=DDB_G0275951\|UniProtKB=Q553A0	Dictyostelium discoideum	O
DICDI\|dictyBase=DDB_G0270160\|UniProtKB=Q55C93	Dictyostelium discoideum	O
DICDI\|dictyBase=DDB_G0285587\|UniProtKB=Q54N01	Dictyostelium discoideum	O
DICPU\|EnsemblGenome=DICPUDRAFT_82882\|UniProtKB=F0ZXX1	Dictyostelium purpureum	O
DICPU\|EnsemblGenome=DICPUDRAFT_92831\|UniProtKB=F0ZXX3	Dictyostelium purpureum	O
THAPS\|EnsemblGenome=THAPSDRAFT_17961\|UniProtKB=B8CB69	Thalassiosira pseudonana	O
PHYRM\|Gene=H3GV46_PHYRM\|UniProtKB=H3GV46	Phytophthora ramorum	O
CHLRE\|EnsemblGenome=CHLRE_07g337700v5\|UniProtKB=A0A2K3DKC0	Chlamydomonas reinhardtii	O
CHLRE\|EnsemblGenome=CHLRE_06g252401v5\|UniProtKB=A8HYH9	Chlamydomonas reinhardtii	O
AMBTC\|EnsemblGenome=AMTR_s00002p00271010\|UniProtKB=W1P1K7	Amborella trichopoda	O
ZOSMR\|Gene_ORFName=ZOSMA_9G00690\|UniProtKB=A0A0K9NGR4	Zostera marina	O
ZOSMR\|Gene_ORFName=ZOSMA_9G00700\|UniProtKB=A0A0K9NIP8	Zostera marina	O
MUSAM\|EnsemblGenome=Ma08_g20480\|UniProtKB=A0A804K8U1	Musa acuminata subsp. malaccensis	O
SETIT\|EnsemblGenome=SETIT_038585mg\|UniProtKB=K4AI76	Setaria italica	O
SORBI\|EnsemblGenome=SORBI_3001G057900\|UniProtKB=C5WX80	Sorghum bicolor	O
ORYSJ\|EnsemblGenome=Os07g0177200\|UniProtKB=Q6ZEY7	Oryza sativa	O
BRADI\|EnsemblGenome=BRADI_1g55660v3\|UniProtKB=I1H359	Brachypodium distachyon	O
ORYSJ\|EnsemblGenome=Os03g0795600\|UniProtKB=Q6F386	Oryza sativa	O
BRADI\|EnsemblGenome=BRADI_1g05570v3\|UniProtKB=I1GM87	Brachypodium distachyon	O
HORVV\|EnsemblGenome=HORVU.MOREX.r3.5HG0518970\|UniProtKB=A0A287SLC8	Hordeum vulgare subsp. vulgare	O
WHEAT\|EnsemblGenome=TraesCS5D02G444900\|UniProtKB=A0A3B6N0T0	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS5A02G437700\|UniProtKB=A0A3B6KR00	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS5B02G440500\|UniProtKB=A0A3B6LTJ7	Triticum aestivum	O
MUSAM\|EnsemblGenome=Ma09_g10260\|UniProtKB=A0A804KI15	Musa acuminata subsp. malaccensis	O
SETIT\|EnsemblGenome=SETIT_026757mg\|UniProtKB=K3ZJK4	Setaria italica	O
SORBI\|EnsemblGenome=SORBI_3005G166100\|UniProtKB=C5Y5A4	Sorghum bicolor	O
MAIZE\|EnsemblGenome=Zm00001eb200530\|UniProtKB=K7UNR6	Zea mays	O
ORYSJ\|EnsemblGenome=Os11g0581900\|UniProtKB=Q2R230	Oryza sativa	O
BRADI\|EnsemblGenome=BRADI_4g15160v3\|UniProtKB=I1IKT9	Brachypodium distachyon	O
MUSAM\|EnsemblGenome=Ma01_g07640\|UniProtKB=A0A804HRE3	Musa acuminata subsp. malaccensis	O
MUSAM\|EnsemblGenome=Ma01_g21270\|UniProtKB=A0A804HWQ2	Musa acuminata subsp. malaccensis	O
MUSAM\|EnsemblGenome=Ma07_g27190\|UniProtKB=A0A804K0E3	Musa acuminata subsp. malaccensis	O
SETIT\|EnsemblGenome=SETIT_037166mg\|UniProtKB=K4AE59	Setaria italica	O
SORBI\|EnsemblGenome=SORBI_3005G071401\|UniProtKB=A0A1Z5RH64	Sorghum bicolor	O
SORBI\|EnsemblGenome=SORBI_3001G092600\|UniProtKB=C5X172	Sorghum bicolor	O
MAIZE\|EnsemblGenome=Zm00001eb215030\|UniProtKB=A0A1D6GHU9	Zea mays	O
MAIZE\|EnsemblGenome=Zm00001eb057010\|UniProtKB=A0A1D6L3W8	Zea mays	O
ORYSJ\|EnsemblGenome=Os03g0745600\|UniProtKB=B7F9F6	Oryza sativa	O
BRADI\|EnsemblGenome=BRADI_1g08870v3\|UniProtKB=I1GND6	Brachypodium distachyon	O
HORVV\|EnsemblGenome=HORVU.MOREX.r3.5HG0455590\|UniProtKB=A0A8I6XAA9	Hordeum vulgare subsp. vulgare	O
HORVV\|EnsemblGenome=HORVU.MOREX.r3.5HG0510030\|UniProtKB=A0A8I6XSH3	Hordeum vulgare subsp. vulgare	O
WHEAT\|EnsemblGenome=TraesCS5B02G388900\|UniProtKB=W5FYB4	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS5A02G384100\|UniProtKB=A0A3B6KPA3	Triticum aestivum	O
NELNU\|Gene=LOC104596610\|UniProtKB=A0A1U7ZRS3	Nelumbo nucifera	O
SPIOL\|Gene=LOC110783460\|UniProtKB=A0A9R0I625	Spinacia oleracea	O
LACSA\|Gene_ORFName=LSAT_V11C800412430\|UniProtKB=A0A9R1UV91	Lactuca sativa	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a012728mg\|UniProtKB=A0A022RKD4	Erythranthe guttata	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a012770mg\|UniProtKB=A0A022S2F0	Erythranthe guttata	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a019280mg\|UniProtKB=A0A022QSQ4	Erythranthe guttata	O
ERYGU\|Gene_ORFName=MIMGU_mgv11b010581mg\|UniProtKB=A0A022S579	Erythranthe guttata	O
CAPAN\|EnsemblGenome=T459_18499\|UniProtKB=A0A1U8H6V4	Capsicum annuum	O
SOLTU\|EnsemblGenome=PGSC0003DMG400020137\|UniProtKB=M1BSE1	Solanum tuberosum	O
SOLLC\|EnsemblGenome=Solyc06g084130.3\|UniProtKB=A0A3Q7H5A5	Solanum lycopersicum	O
TOBAC\|Gene=LOC107776210\|UniProtKB=A0A1S3YHH1	Nicotiana tabacum	O
TOBAC\|Gene=LOC107810984\|UniProtKB=A0A1S4BQZ4	Nicotiana tabacum	O
VITVI\|EnsemblGenome=Vitvi07g00087\|UniProtKB=D7TE15	Vitis vinifera	O
BRANA\|Gene_ORFName=DARMORV10_A04P00110.1\|UniProtKB=A0A078IV04	Brassica napus	O
BRACM\|EnsemblGenome=Bra040445\|UniProtKB=M4FH67	Brassica campestris	O
ARATH\|TAIR=locus=2077309\|UniProtKB=Q9M1V9	Arabidopsis thaliana	O
POPTR\|EnsemblGenome=POPTR_005G213900v3\|UniProtKB=B9H8C6	Populus trichocarpa	O
MANES\|EnsemblGenome=MANES_05G080700\|UniProtKB=A0A2C9VWS9	Manihot esculenta	O
RICCO\|Gene_ORFName=RCOM_0646050\|UniProtKB=B9SFF9	Ricinus communis	O
EUCGR\|EnsemblGenome=EUGRSUZ_H01638\|UniProtKB=A0A059AYG8	Eucalyptus grandis	O
EUCGR\|EnsemblGenome=EUGRSUZ_K03330\|UniProtKB=A0A059A6X9	Eucalyptus grandis	O
BRANA\|Gene_ORFName=DARMORV10_C01P28510.1\|UniProtKB=A0A078FP74	Brassica napus	O
BRANA\|Gene_ORFName=DARMORV10_A01P22880.1\|UniProtKB=D2CVS1	Brassica napus	O
BRACM\|EnsemblGenome=Bra036891\|UniProtKB=M4F731	Brassica campestris	O
ARATH\|TAIR=locus=2130170\|UniProtKB=F4JIE8	Arabidopsis thaliana	O
GOSHI\|Gene=LOC107951221\|UniProtKB=A0A1U8NUC4	Gossypium hirsutum	O
GOSHI\|Gene=LOC107914862\|UniProtKB=A0A1U8KA83	Gossypium hirsutum	O
BRANA\|Gene_ORFName=GSBRNA2T00092257001\|UniProtKB=A0A078II46	Brassica napus	O
BRACM\|EnsemblGenome=Bra031411\|UniProtKB=A0A3P5Z9L1	Brassica campestris	O
BRACM\|EnsemblGenome=Bra018534\|UniProtKB=M4DPU3	Brassica campestris	O
ARATH\|TAIR=locus=2132318\|UniProtKB=Q9ZQX7	Arabidopsis thaliana	O
BRANA\|Gene_ORFName=DARMORV10_C05P01690.1\|UniProtKB=A0A078FZE8	Brassica napus	O
BRACM\|EnsemblGenome=Bra033373\|UniProtKB=A0A3P6CEC2	Brassica campestris	O
ARATH\|TAIR=locus=2007509\|UniProtKB=Q9SA63	Arabidopsis thaliana	O
GOSHI\|Gene=LOC107936473\|UniProtKB=A0A1U8MCX7	Gossypium hirsutum	O
GOSHI\|Gene=LOC107940222\|UniProtKB=A0A1U8MQQ3	Gossypium hirsutum	O
GOSHI\|Gene=LOC107960958\|UniProtKB=A0A1U8PNG3	Gossypium hirsutum	O
THECC\|EnsemblGenome=TCM_004291\|UniProtKB=A0A061DQL7	Theobroma cacao	O
CITSI\|Gene_ORFName=CISIN_1g026195mg\|UniProtKB=A0A067FMM1	Citrus sinensis	O
POPTR\|EnsemblGenome=POPTR_005G214000v3\|UniProtKB=B9H8C7	Populus trichocarpa	O
POPTR\|EnsemblGenome=POPTR_002G049000v3\|UniProtKB=B9GSJ5	Populus trichocarpa	O
MANES\|EnsemblGenome=MANES_05G080800\|UniProtKB=A0A2C9VVR6	Manihot esculenta	O
MANES\|EnsemblGenome=MANES_01G204600\|UniProtKB=A0A2C9WMM2	Manihot esculenta	O
RICCO\|Gene_ORFName=RCOM_0646040\|UniProtKB=B9SFF8	Ricinus communis	O
RICCO\|Gene_ORFName=RCOM_0646030\|UniProtKB=B9SFF7	Ricinus communis	O
SOYBN\|EnsemblGenome=GLYMA_10G272600\|UniProtKB=I1LEV2	Glycine max	O
SOYBN\|EnsemblGenome=GLYMA_20G117300\|UniProtKB=I1NFL0	Glycine max	O
MEDTR\|EnsemblGenome=MTR_1g110390\|UniProtKB=G7ICA0	Medicago truncatula	O
SOYBN\|EnsemblGenome=GLYMA_19G159300\|UniProtKB=K7MYM2	Glycine max	O
SOYBN\|EnsemblGenome=GLYMA_02G144700\|UniProtKB=A0A0R0L253	Glycine max	O
SOYBN\|EnsemblGenome=GLYMA_10G029400\|UniProtKB=C6T4L3	Glycine max	O
MEDTR\|EnsemblGenome=MTR_1g071430\|UniProtKB=G7IDK0	Medicago truncatula	O
JUGRE\|EnsemblGenome=Jr12_15530\|UniProtKB=A0A2I4G5L8	Juglans regia	O
CUCSA\|EnsemblGenome=Csa_2G222100\|UniProtKB=A0A0A0LLN6	Cucumis sativus	O
JUGRE\|EnsemblGenome=Jr08_14030\|UniProtKB=A0A2I4E5M7	Juglans regia	O
JUGRE\|Gene=LOC109018640\|UniProtKB=A0A2I4HJI9	Juglans regia	O
CUCSA\|EnsemblGenome=Csa_2G223750\|UniProtKB=A0A0A0LKU5	Cucumis sativus	O
PRUPE\|EnsemblGenome=PRUPE_8G235700\|UniProtKB=M5VKA6	Prunus persica	O
SELML\|EnsemblGenome=SELMODRAFT_430816\|UniProtKB=D8TAM1	Selaginella moellendorffii	O
SELML\|EnsemblGenome=SELMODRAFT_84281\|UniProtKB=D8R4M6	Selaginella moellendorffii	O
SELML\|EnsemblGenome=SELMODRAFT_95705\|UniProtKB=D8RK20	Selaginella moellendorffii	O
SELML\|EnsemblGenome=SELMODRAFT_88443\|UniProtKB=D8R9A6	Selaginella moellendorffii	O
SELML\|EnsemblGenome=SELMODRAFT_150230\|UniProtKB=D8RV78	Selaginella moellendorffii	O
PHYPA\|EnsemblGenome=Pp3c4_29330\|UniProtKB=A9SQ31	Physcomitrella patens	O
PHYPA\|EnsemblGenome=Pp3c9_13460\|UniProtKB=A0A2K1K328	Physcomitrella patens	O
PHYPA\|EnsemblGenome=Pp3c15_13530\|UniProtKB=A0A2K1JD14	Physcomitrella patens	O
PHYPA\|EnsemblGenome=Pp3c3_15530\|UniProtKB=A0A2K1KUL9	Physcomitrella patens	O
MARPO\|EnsemblGenome=MARPO_0093s0068\|UniProtKB=A0A2R6WGM3	Marchantia polymorpha	O
KLENI\|Gene_ORFName=KFL_003980150\|UniProtKB=A0A1Y1IAY3	Klebsormidium nitens	O
KLENI\|Gene_ORFName=KFL_001530050\|UniProtKB=A0A1Y1I0S1	Klebsormidium nitens	O
STRR6\|EnsemblGenome=spr1787\|UniProtKB=Q8DNE4	Streptococcus pneumoniae	O
STAA8\|EnsemblGenome=SAOUHSC_00672\|UniProtKB=Q2G0D3	Staphylococcus aureus	O
LISMO\|Gene=Q8Y569_LISMO\|UniProtKB=Q8Y569	Listeria monocytogenes	O
BACSU\|EnsemblGenome=BSU_07200\|UniProtKB=O31539	Bacillus subtilis	O
DEIRA\|EnsemblGenome=DR_0893\|UniProtKB=Q9RVX8	Deinococcus radiodurans	O
GLOVI\|Gene=Q7NPB6_GLOVI\|UniProtKB=Q7NPB6	Gloeobacter violaceus	O
SYNY3\|Gene=P73793_SYNY3\|UniProtKB=P73793	Synechocystis	O
CHLTR\|EnsemblGenome=CT_819\|UniProtKB=O84826	Chlamydia trachomatis	O
RHOBA\|EnsemblGenome=RB3430\|UniProtKB=Q7UU92	Rhodopirellula baltica	O
FUSNN\|EnsemblGenome=FN0866\|UniProtKB=Q8RF52	Fusobacterium nucleatum	O
HELPY\|EnsemblGenome=HP_0920\|UniProtKB=O25578	Helicobacter pylori	O
NEIMB\|EnsemblGenome=NMB2020\|UniProtKB=P63702	Neisseria meningitidis serogroup b	O
HAEIN\|EnsemblGenome=HI_0044\|UniProtKB=P44477	Haemophilus influenzae	O
SHEON\|Gene_OrderedLocusName=SO_2375\|UniProtKB=Q8EEK6	Shewanella oneidensis	O
VIBCH\|EnsemblGenome=VC_1358\|UniProtKB=Q9KSA1	Vibrio cholerae	O
PSEAE\|EnsemblGenome=PA2604\|UniProtKB=Q03268	Pseudomonas aeruginosa	O
SALTY\|EnsemblGenome=STM1085\|UniProtKB=Q8ZQ61	Salmonella typhimurium	O
ECOLI\|EnsemblGenome=BAA35735\|UniProtKB=P0AAC6	Escherichia coli	O
BRADU\|Gene=Q89X53_BRADU\|UniProtKB=Q89X53	Bradyrhizobium diazoefficiens	O
BRADU\|Gene=Q89DW5_BRADU\|UniProtKB=Q89DW5	Bradyrhizobium diazoefficiens	O
COXBU\|EnsemblGenome=CBU_0214\|UniProtKB=Q83EU5	Coxiella burnetii	O
SHEON\|Gene_OrderedLocusName=SO_0136\|UniProtKB=Q8EKF7	Shewanella oneidensis	O
PSEAE\|EnsemblGenome=PA1190\|UniProtKB=Q9I4E8	Pseudomonas aeruginosa	O
SALTY\|EnsemblGenome=STM0807\|UniProtKB=Q8ZQP9	Salmonella typhimurium	O
YERPE\|Gene_OrderedLocusName=YPO1163\|UniProtKB=A0A2U2GWC9	Yersinia pestis	O
ECOLI\|EnsemblGenome=BAA35444\|UniProtKB=P0AAC4	Escherichia coli	O