PANTHER Gene Information

Gene Symbol(s):

GMPPA

Organism:

Equus caballus

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Tree

Gene Name:

GDP-mannose pyrophosphorylase A

Gene ID:

ENSECAG00000011565.4

Protein ID:

F7BEM5

Persistent Id:

PTN001475477

Alternate Ids:

194211348(GI)	XP_005610691(refSeq)
ENSECAP00000009743(Ensembl_PRO)	100059471(GeneID)
F7BEM5(Synonym)	ENSECAT00000012341(Ensembl_TRS)
F7BEM5_HORSE(UniProtKB-ID)	ENSECAG00000011565(Ensembl)
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PANTHER Classification

PANTHER Family:	SUGAR-1-PHOSPHATE GUANYL TRANSFERASE (PTHR22572)
PANTHER Subfamily:	MANNOSE-1-PHOSPHATE GUANYLTRANSFERASE ALPHA (PTHR22572:SF104)
PANTHER GO-slim Molecular Function:	nucleotidyltransferase activity
PANTHER GO-slim Biological Process:
PANTHER GO-slim Cellular Component:	cytoplasm
PANTHER protein class:	transferase

Pathway Categories:

Mannose metabolism

Mannose 1-P guanylyl transferase

GENE ONTOLOGY DATABASE ANNOTATIONS

Click to Expand...

GO MF Complete:	enzyme inhibitor activity, transferase activity, enzyme binding
GO BP Complete:	negative regulation of small molecule metabolic process, negative regulation of phosphate metabolic process, telencephalon development, glycoprotein metabolic process, GDP-mannose biosynthetic process, negative regulation of biosynthetic process, neuromuscular process, protein glycosylation, skeletal muscle organ development, biosynthetic process, muscle organ morphogenesis, negative regulation of nucleobase-containing compound metabolic process, motor behavior, cognition, GDP-mannose metabolic process, neuron apoptotic process
GO CC Complete:	cytoplasm, GDP-mannose pyrophosphorylase complex

REACTOME DATABASE ANNOTATIONS Click to Expand...

Reactome Pathways:

Orthologs

ID	Organism	Type
HUMAN\|HGNC=22923\|UniProtKB=Q96IJ6	Homo sapiens	LDO
PANTR\|Ensembl=ENSPTRG00000012959.6\|UniProtKB=H2QJH1	Pan troglodytes	LDO
GORGO\|Ensembl=ENSGGOG00000002725.3\|UniProtKB=G3SHF2	Gorilla gorilla gorilla	LDO
MACMU\|Ensembl=ENSMMUG00000017938.4\|UniProtKB=A0A1D5QED6	Macaca mulatta	LDO
MOUSE\|MGI=MGI=1916330\|UniProtKB=Q922H4	Mus musculus	LDO
RAT\|RGD=1560644\|UniProtKB=Q5XIC1	Rattus norvegicus	LDO
BOVIN\|Ensembl=ENSBTAG00000002995.5\|UniProtKB=E1BEN4	Bos taurus	LDO
PIG\|Ensembl=ENSSSCG00000023935.4\|UniProtKB=A0A140TAL0	Sus scrofa	LDO
PIG\|Gene=GMPPA\|UniProtKB=I3LUP1	Sus scrofa	LDO
FELCA\|Ensembl=ENSFCAG00000022252.4\|UniProtKB=M3WU81	Felis catus	LDO
CANLF\|Ensembl=ENSCAFG00845030366.1\|UniProtKB=A0A8I3Q9B9	Canis lupus familiaris	LDO
MONDO\|Ensembl=ENSMODG00000015078.4\|UniProtKB=F7F3V5	Monodelphis domestica	LDO
ORNAN\|Ensembl=ENSOANG00000042671.1\|UniProtKB=A0A6I8NGY9	Ornithorhynchus anatinus	LDO
CHICK\|Ensembl=ENSGALG00010018209.1\|UniProtKB=F1NYN9	Gallus gallus	LDO
ANOCA\|Ensembl=ENSACAG00000038354.1\|UniProtKB=A0A803T389	Anolis carolinensis	O
ANOCA\|Ensembl=ENSACAG00000014108.4\|UniProtKB=A0A803SV96	Anolis carolinensis	LDO
XENLA\|Gene=gmppa.S\|UniProtKB=A0A8J0TXQ2	Xenopus laevis	O
XENLA\|Gene=gmppa-a\|UniProtKB=Q6DKE9	Xenopus laevis	O
XENLA\|Gene=gmppa.L\|UniProtKB=A0A1L8EVM9	Xenopus laevis	LDO
XENLA\|Gene=gmppa-b\|UniProtKB=Q66KG5	Xenopus laevis	LDO
XENTR\|Xenbase=XB-GENE-984239\|UniProtKB=A0A5S6N5I2	Xenopus tropicalis	LDO
XENTR\|Gene=gmppa\|UniProtKB=Q0VFM6	Xenopus tropicalis	LDO
ORYLA\|Ensembl=ENSORLG00000020513.2\|UniProtKB=A0A3B3HLK9	Oryzias latipes	LDO
DANRE\|ZFIN=ZDB-GENE-040426-1550\|UniProtKB=Q7SXP8	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-040704-37\|UniProtKB=Q6GMK8	Danio rerio	LDO
LEPOC\|Ensembl=ENSLOCG00000000759.1\|UniProtKB=W5LXI7	lepisosteus oculatus	LDO
CIOIN\|Ensembl=ENSCING00000006662.3\|UniProtKB=F6Q0G0	Ciona intestinalis	O
CIOIN\|Ensembl=ENSCING00000011553.2\|UniProtKB=F6TC27	Ciona intestinalis	LDO
BRAFL\|Gene=LOC118409757\|UniProtKB=A0A9J7MGG7	Branchiostoma floridae	LDO
STRPU\|EnsemblGenome=LOC576782\|UniProtKB=A0A7M7GEW4	Strongylocentrotus purpuratus	LDO
CAEEL\|WormBase=WBGene00021628\|UniProtKB=Q9N4V3	Caenorhabditis elegans	LDO
CAEBR\|Gene_ORFName=CBG12186\|UniProtKB=A8XF15	Caenorhabditis briggsae	LDO
PRIPA\|EnsemblGenome=WBGene00099278\|UniProtKB=A0A2A6BVA3	Pristionchus pacificus	LDO
DROME\|FlyBase=FBgn0034035\|UniProtKB=Q8SXU3	Drosophila melanogaster	LDO
ANOGA\|EnsemblGenome=AGAP011723\|UniProtKB=Q5TML0	Anopheles gambiae	LDO
TRICA\|EnsemblGenome=TC033614\|UniProtKB=A0A139WFJ8	Tribolium castaneum	LDO
DAPPU\|EnsemblGenome=DAPPUDRAFT_191301\|UniProtKB=E9FUH9	Daphnia pulex	LDO
IXOSC\|EnsemblGenome=ISCW015357\|UniProtKB=B7QN92	Ixodes scapularis	LDO
HELRO\|EnsemblGenome=HelroG185698\|UniProtKB=T1FN59	helobdella robusta	LDO
NEMVE\|EnsemblGenome=NEMVEDRAFT_v1g240817\|UniProtKB=A7RUP7	Nematostella vectensis	LDO
TRIAD\|EnsemblGenome=TriadG22620\|UniProtKB=B3RS02	Trichoplax adhaerens	LDO
MONBE\|EnsemblGenome=MONBRDRAFT_33433\|UniProtKB=A9V5D1	Monosiga brevicollis	LDO
CANAL\|CGD=CAL0000185249\|UniProtKB=Q5AL34	Candida albicans	LDO
YARLI\|EnsemblGenome=YALI0_E15125g\|UniProtKB=Q6C5U4	Yarrowia lipolytica	LDO
EMENI\|EnsemblGenome=ANIA_01911\|UniProtKB=C8VKT1	Emericella nidulans	LDO
ASPFU\|EnsemblGenome=AFUA_6G07620\|UniProtKB=Q4WN49	Neosartorya fumigata	LDO
NEUCR\|EnsemblGenome=NCU05937\|UniProtKB=Q7S2E1	Neurospora crassa	LDO
SCLS1\|EnsemblGenome=SS1G_13453\|UniProtKB=A7F773	Sclerotinia sclerotiorum	LDO
PHANO\|EnsemblGenome=SNOG_06386\|UniProtKB=Q0UPC8	Phaeosphaeria nodorum	LDO
SCHPO\|PomBase=SPBC13G1.02\|UniProtKB=O60064	Schizosaccharomyces pombe	LDO
SCHJY\|EnsemblGenome=SJAG_00890\|UniProtKB=B6JWW5	Schizosaccharomyces japonicus	LDO
CRYNJ\|EnsemblGenome=CNG02580\|UniProtKB=Q5KDW3	Cryptococcus neoformans	LDO
PUCGT\|EnsemblGenome=PGTG_17414\|UniProtKB=E3L628	Puccinia graminis	LDO
BATDJ\|Gene_ORFName=BATDEDRAFT_33853\|UniProtKB=F4PEM3	Batrachochytrium dendrobatidis	LDO
DICDI\|dictyBase=DDB_G0271858\|UniProtKB=Q86HG0	Dictyostelium discoideum	LDO
DICPU\|EnsemblGenome=DICPUDRAFT_159905\|UniProtKB=F1A590	Dictyostelium purpureum	LDO
PARTE\|EnsemblGenome=GSPATT00000654001\|UniProtKB=A0CJH6	Paramecium tetraurelia	O
PARTE\|EnsemblGenome=GSPATT00032380001\|UniProtKB=A0BUD1	Paramecium tetraurelia	LDO
PHYRM\|Gene=H3GFQ5_PHYRM\|UniProtKB=H3GFQ5	Phytophthora ramorum	LDO
AMBTC\|EnsemblGenome=AMTR_s00058p00031570\|UniProtKB=W1PH93	Amborella trichopoda	LDO
ZOSMR\|Gene_ORFName=ZOSMA_87G00020\|UniProtKB=A0A0K9NMQ8	Zostera marina	LDO
MUSAM\|EnsemblGenome=Ma03_g31450\|UniProtKB=A0A804IIG0	Musa acuminata subsp. malaccensis	LDO
MUSAM\|EnsemblGenome=Ma10_g18160\|UniProtKB=A0A804KXJ4	Musa acuminata subsp. malaccensis	O
MUSAM\|EnsemblGenome=Ma07_g27920\|UniProtKB=A0A804K0P2	Musa acuminata subsp. malaccensis	LDO
SETIT\|EnsemblGenome=SETIT_035885mg\|UniProtKB=K4AAH8	Setaria italica	LDO
SORBI\|EnsemblGenome=SORBI_3001G462200\|UniProtKB=C5WTQ1	Sorghum bicolor	LDO
MAIZE\|EnsemblGenome=Zm00001eb400530\|UniProtKB=A0A804RBR8	Zea mays	LDO
MAIZE\|EnsemblGenome=Zm00001eb007180\|UniProtKB=C0PJV3	Zea mays	LDO
ORYSJ\|EnsemblGenome=Os03g0208900\|UniProtKB=Q10Q61	Oryza sativa	LDO
BRADI\|EnsemblGenome=BRADI_1g70620v3\|UniProtKB=I1H8B7	Brachypodium distachyon	LDO
HORVV\|EnsemblGenome=HORVU.MOREX.r3.4HG0397900\|UniProtKB=F2DTF1	Hordeum vulgare subsp. vulgare	LDO
WHEAT\|EnsemblGenome=TraesCS4B02G253000\|UniProtKB=A0A3B6ITY1	Triticum aestivum	LDO
WHEAT\|EnsemblGenome=TraesCS4A02G051400\|UniProtKB=A0A3B6HNC2	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS4D02G253100\|UniProtKB=A0A3B6JKE9	Triticum aestivum	LDO
NELNU\|Gene=LOC104607977\|UniProtKB=A0A1U8B7Y0	Nelumbo nucifera	O
NELNU\|Gene=LOC104596039\|UniProtKB=A0A1U7ZML7	Nelumbo nucifera	LDO
SPIOL\|Gene=LOC110786830\|UniProtKB=A0A9R0IDM9	Spinacia oleracea	O
HELAN\|Gene_ORFName=HanXRQr2_Chr13g0570981\|UniProtKB=A0A9K3EGK9	Helianthus annuus	O
HELAN\|EnsemblGenome=HanXRQr2_Chr09g0402361\|UniProtKB=A0A251TXR8	Helianthus annuus	O
LACSA\|Gene_ORFName=LSAT_V11C400225970\|UniProtKB=A0A9R1XGC5	Lactuca sativa	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a007245mg\|UniProtKB=A0A022RXA1	Erythranthe guttata	O
TOBAC\|Gene=LOC107767406\|UniProtKB=A0A1S3XQE1	Nicotiana tabacum	O
TOBAC\|Gene=LOC107767184\|UniProtKB=A0A1S3XNP2	Nicotiana tabacum	O
VITVI\|Gene_OrderedLocusName=VIT_14s0066g02400\|UniProtKB=D7TWZ7	Vitis vinifera	O
EUCGR\|EnsemblGenome=EUGRSUZ_I01880\|UniProtKB=A0A059ARS2	Eucalyptus grandis	O
BRANA\|Gene_ORFName=DARMORV10_A04P15650.1\|UniProtKB=A0A078FS38	Brassica napus	O
BRACM\|EnsemblGenome=Bra030345\|UniProtKB=M4ENH5	Brassica campestris	O
ARATH\|TAIR=locus=2049188\|UniProtKB=F4IFA4	Arabidopsis thaliana	O
GOSHI\|Gene=LOC107958596\|UniProtKB=A0A1U8PI64	Gossypium hirsutum	O
GOSHI\|Gene=LOC107955565\|UniProtKB=A0A1U8P9C3	Gossypium hirsutum	O
THECC\|EnsemblGenome=TCM_017958\|UniProtKB=A0A061EEA3	Theobroma cacao	O
CITSI\|Gene_ORFName=CISIN_1g015225mg\|UniProtKB=A0A067FS72	Citrus sinensis	O
RICCO\|Gene_ORFName=RCOM_0852750\|UniProtKB=B9RUH2	Ricinus communis	O
SOYBN\|EnsemblGenome=GLYMA_19G050100\|UniProtKB=I1N6T7	Glycine max	O
MEDTR\|EnsemblGenome=MTR_6g008840\|UniProtKB=G7KJH0	Medicago truncatula	O
JUGRE\|EnsemblGenome=Jr01_03740\|UniProtKB=A0A2I4EUR2	Juglans regia	O
CUCSA\|EnsemblGenome=Csa_3G829100\|UniProtKB=A0A0A0LC79	Cucumis sativus	O
PRUPE\|EnsemblGenome=PRUPE_3G276900\|UniProtKB=M5XC41	Prunus persica	O
SPIOL\|Gene=LOC110776900\|UniProtKB=A0A9R0JK09	Spinacia oleracea	LDO
HELAN\|EnsemblGenome=HanXRQr2_Chr16g0730321\|UniProtKB=A0A251UWM6	Helianthus annuus	O
HELAN\|EnsemblGenome=HanXRQr2_Chr09g0410001\|UniProtKB=A0A251TZ94	Helianthus annuus	LDO
LACSA\|Gene_ORFName=LSAT_V11C400186690\|UniProtKB=A0A9R1XGZ1	Lactuca sativa	LDO
ERYGU\|Gene_ORFName=MIMGU_mgv1a023858mg\|UniProtKB=A0A022QGA2	Erythranthe guttata	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a022325mg\|UniProtKB=A0A022QXS9	Erythranthe guttata	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a007207mg\|UniProtKB=A0A022RRB0	Erythranthe guttata	LDO
CAPAN\|EnsemblGenome=T459_29880\|UniProtKB=A0A1U8FWM6	Capsicum annuum	LDO
SOLTU\|EnsemblGenome=PGSC0003DMG402024581\|UniProtKB=M1CA88	Solanum tuberosum	LDO
SOLLC\|EnsemblGenome=Solyc03g113790.3\|UniProtKB=A0A3Q7FQ28	Solanum lycopersicum	LDO
TOBAC\|Gene=LOC107796939\|UniProtKB=A0A1S4AF29	Nicotiana tabacum	O
TOBAC\|Gene=LOC107773227\|UniProtKB=A0A1S3Y7B8	Nicotiana tabacum	O
TOBAC\|Gene=LOC107806352\|UniProtKB=A0A1S4BAT4	Nicotiana tabacum	LDO
VITVI\|EnsemblGenome=Vitvi17g00738\|UniProtKB=D7SHY4	Vitis vinifera	LDO
EUCGR\|EnsemblGenome=EUGRSUZ_K02882\|UniProtKB=A0A059A685	Eucalyptus grandis	LDO
BRANA\|Gene_ORFName=GSBRNA2T00083141001\|UniProtKB=A0A078I9H5	Brassica napus	O
BRANA\|Gene_ORFName=GSBRNA2T00063780001\|UniProtKB=A0A078HFW4	Brassica napus	O
BRACM\|EnsemblGenome=Bra008171\|UniProtKB=M4CVC4	Brassica campestris	O
BRANA\|Gene_ORFName=DARMORV10_A07P40540.1\|UniProtKB=A0A078I7M5	Brassica napus	LDO
BRACM\|EnsemblGenome=Bra015883\|UniProtKB=M4DHA6	Brassica campestris	LDO
ARATH\|TAIR=locus=2027201\|UniProtKB=Q9C9P3	Arabidopsis thaliana	LDO
GOSHI\|Gene=LOC107926700\|UniProtKB=A0A1U8LEQ8	Gossypium hirsutum	O
GOSHI\|Gene=LOC107953544\|UniProtKB=A0A1U8P1Z9	Gossypium hirsutum	LDO
THECC\|EnsemblGenome=TCM_011970\|UniProtKB=A0A061FU19	Theobroma cacao	LDO
CITSI\|Gene_ORFName=CISIN_1g0152591mg\|UniProtKB=A0A067G279	Citrus sinensis	LDO
POPTR\|EnsemblGenome=POPTR_012G075500v3\|UniProtKB=B9I350	Populus trichocarpa	O
POPTR\|EnsemblGenome=POPTR_015G070500v3\|UniProtKB=B9IEL3	Populus trichocarpa	LDO
MANES\|EnsemblGenome=MANES_14G070700\|UniProtKB=A0A2C9UJJ5	Manihot esculenta	O
MANES\|EnsemblGenome=MANES_06G100100\|UniProtKB=A0A2C9VPG9	Manihot esculenta	LDO
RICCO\|Gene_ORFName=RCOM_1512730\|UniProtKB=B9R8Y9	Ricinus communis	LDO
SOYBN\|EnsemblGenome=GLYMA_04G178400\|UniProtKB=I1JX24	Glycine max	LDO
SOYBN\|EnsemblGenome=GLYMA_06G186400\|UniProtKB=K7KVU9	Glycine max	LDO
MEDTR\|EnsemblGenome=MTR_3g078570\|UniProtKB=A0A072UZU5	Medicago truncatula	LDO
JUGRE\|EnsemblGenome=Jr02_04170\|UniProtKB=A0A2I4FIR6	Juglans regia	LDO
JUGRE\|EnsemblGenome=Jr02_04160\|UniProtKB=A0A2I4FIR7	Juglans regia	LDO
CUCSA\|EnsemblGenome=Csa_1G006280\|UniProtKB=A0A0A0LP89	Cucumis sativus	LDO
PRUPE\|EnsemblGenome=PRUPE_5G179900\|UniProtKB=A0A251PBQ5	Prunus persica	LDO
SELML\|EnsemblGenome=SELMODRAFT_182999\|UniProtKB=D8SV91	Selaginella moellendorffii	LDO
SELML\|EnsemblGenome=SELMODRAFT_104991\|UniProtKB=D8RZ45	Selaginella moellendorffii	LDO
PHYPA\|EnsemblGenome=Pp3c20_18250\|UniProtKB=A0A2K1IVP9	Physcomitrella patens	LDO
PHYPA\|EnsemblGenome=Pp3c24_4010\|UniProtKB=A0A2K1IFE3	Physcomitrella patens	O
PHYPA\|EnsemblGenome=Pp3c8_6490\|UniProtKB=A9RGJ6	Physcomitrella patens	O
MARPO\|EnsemblGenome=MARPO_0163s0013\|UniProtKB=A0A2R6W3M5	Marchantia polymorpha	LDO
KLENI\|Gene_ORFName=KFL_007060020\|UniProtKB=A0A1Y1IJ89	Klebsormidium nitens	LDO
NITMS\|EnsemblGenome=Nmar_0424\|UniProtKB=A9A5C1	Nitrosopumilus maritimus	O
NITMS\|EnsemblGenome=Nmar_0843\|UniProtKB=A9A5M8	Nitrosopumilus maritimus	O
SACS2\|EnsemblGenome=SSO0317\|UniProtKB=Q980H9	Sulfolobus solfataricus	O
PYRAE\|EnsemblGenome=PAE3554\|UniProtKB=Q8ZSW5	Pyrobaculum aerophilum	O
SACS2\|EnsemblGenome=SSO0381\|UniProtKB=Q980D4	Sulfolobus solfataricus	O
SACS2\|EnsemblGenome=SSO2479\|UniProtKB=Q97VX4	Sulfolobus solfataricus	O
HALSA\|EnsemblGenome=VNG_0390G\|UniProtKB=Q9HS61	Halobacterium salinarum	O
THEKO\|EnsemblGenome=TK1711\|UniProtKB=Q5JJ02	Thermococcus kodakaraensis	O
METAC\|EnsemblGenome=MA_3140\|UniProtKB=Q8TL99	Methanosarcina acetivorans	O
THEKO\|EnsemblGenome=TK0955\|UniProtKB=Q5JI97	Thermococcus kodakaraensis	O
CHLAA\|EnsemblGenome=Caur_2687\|UniProtKB=A9WJI8	Chloroflexus aurantiacus	O
STRCO\|Gene_OrderedLocusName=SCO3039\|UniProtKB=Q9KZK6	Streptomyces coelicolor	O
MYCTU\|Gene_OrderedLocusName=Rv3264c\|UniProtKB=L7N6A5	Mycobacterium tuberculosis	O
CHLAA\|EnsemblGenome=Caur_1473\|UniProtKB=A9WAE2	Chloroflexus aurantiacus	O
CHLAA\|EnsemblGenome=Caur_1759\|UniProtKB=A9WCI5	Chloroflexus aurantiacus	O
CHLAA\|EnsemblGenome=Caur_2960\|UniProtKB=A9WG05	Chloroflexus aurantiacus	O
STRCO\|Gene_OrderedLocusName=SCO1388\|UniProtKB=Q9KZP4	Streptomyces coelicolor	O
BACCR\|EnsemblGenome=BC_4264\|UniProtKB=Q818I9	Bacillus cereus	O
GLOVI\|Gene=Q7NII3_GLOVI\|UniProtKB=Q7NII3	Gloeobacter violaceus	O
GLOVI\|Gene=Q7NF24_GLOVI\|UniProtKB=Q7NF24	Gloeobacter violaceus	O
GLOVI\|Gene=rfbM\|UniProtKB=Q7NJQ0	Gloeobacter violaceus	O
GLOVI\|Gene=Q7NNE0_GLOVI\|UniProtKB=Q7NNE0	Gloeobacter violaceus	O
GLOVI\|Gene=Q7NC95_GLOVI\|UniProtKB=Q7NC95	Gloeobacter violaceus	O
SYNY3\|EnsemblGenome=BAA18379\|UniProtKB=P74285	Synechocystis	O
GLOVI\|Gene=Q7NG32_GLOVI\|UniProtKB=Q7NG32	Gloeobacter violaceus	O
SYNY3\|Gene=P74589_SYNY3\|UniProtKB=P74589	Synechocystis	O
LEPIN\|EnsemblGenome=LA_1581\|UniProtKB=Q8F5T6	Leptospira interrogans	O
LEPIN\|EnsemblGenome=LA_1611\|UniProtKB=Q8F5Q6	Leptospira interrogans	O
AQUAE\|EnsemblGenome=aq_718\|UniProtKB=O66933	Aquifex aeolicus	O
DICTD\|EnsemblGenome=Dtur_1427\|UniProtKB=B8E0W4	Dictyoglomus turgidum	O
GEOSL\|EnsemblGenome=GSU1968\|UniProtKB=Q74B34	Geobacter sulfurreducens	O
GEOSL\|EnsemblGenome=GSU3254\|UniProtKB=Q747L1	Geobacter sulfurreducens	O
BRADU\|Gene=Q89HJ6_BRADU\|UniProtKB=Q89HJ6	Bradyrhizobium diazoefficiens	O