PANTHER Gene Information

Gene Symbol(s):

100779367

Organism:

Glycine max

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Tree

Gene Name:

mannose-1-phosphate guanylyltransferase

Gene ID:

GLYMA_11G223700

Protein ID:

C6TI75

Persistent Id:

PTN001859814

Alternate Ids:

947082254(GI)	NP_001240188(refSeq)
GLYMA_11G223700(Ensembl)	BT097353(EMBL)
C6TI75(Synonym)	KRH31042(EMBL-CDS)
C6TI75_SOYBN(UniProtKB-ID)	100779367(GeneID)
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PANTHER Classification

PANTHER Family:	SUGAR-1-PHOSPHATE GUANYL TRANSFERASE (PTHR22572)
PANTHER Subfamily:	MANNOSE-1-PHOSPHATE GUANYLTRANSFERASE BETA (PTHR22572:SF15)
PANTHER GO-slim Molecular Function:	nucleotidyltransferase activity
PANTHER GO-slim Biological Process:
PANTHER GO-slim Cellular Component:	cytoplasm
PANTHER protein class:	transferase

Pathway Categories:

Mannose metabolism

Mannose 1-P guanylyl transferase

GENE ONTOLOGY DATABASE ANNOTATIONS

Click to Expand...

GO MF Complete:	GTP binding, transferase activity, mannose-1-phosphate guanylyltransferase (GTP) activity
GO BP Complete:	GDP-mannose biosynthetic process, protein glycosylation
GO CC Complete:	cytoplasm

REACTOME DATABASE ANNOTATIONS Click to Expand...

Reactome Pathways:

Orthologs

ID	Organism	Type
HUMAN\|HGNC=22932\|UniProtKB=Q9Y5P6	Homo sapiens	O
PANTR\|Ensembl=ENSPTRG00000014932.5\|UniProtKB=H2QMM2	Pan troglodytes	O
GORGO\|Ensembl=ENSGGOG00000010628.3\|UniProtKB=G3R4Z4	Gorilla gorilla gorilla	O
MACMU\|Ensembl=ENSMMUG00000014584.4\|UniProtKB=F7GRF0	Macaca mulatta	O
MOUSE\|MGI=MGI=2660880\|UniProtKB=Q8BTZ7	Mus musculus	O
RAT\|RGD=1560458\|UniProtKB=D4A746	Rattus norvegicus	O
BOVIN\|Gene=GMPPB\|UniProtKB=Q2YDJ9	Bos taurus	O
BOVIN\|Ensembl=ENSBTAG00000032026.2\|UniProtKB=F1N7H5	Bos taurus	O
PIG\|Ensembl=ENSSSCG00000062528.1\|UniProtKB=A0A8W4F9L7	Sus scrofa	O
PIG\|Gene=GMPPB\|UniProtKB=P0C5I2	Sus scrofa	O
HORSE\|Ensembl=ENSECAG00000019354.3\|UniProtKB=F6VTV5	Equus caballus	O
FELCA\|Ensembl=ENSFCAG00000010676.5\|UniProtKB=M3WDQ0	Felis catus	O
CANLF\|Ensembl=ENSCAFG00845025132.1\|UniProtKB=A0A8I3PK79	Canis lupus familiaris	O
MONDO\|Ensembl=ENSMODG00000012906.3\|UniProtKB=F7GKQ3	Monodelphis domestica	O
ORNAN\|Ensembl=ENSOANG00000050893.1\|UniProtKB=A0A6I8PL06	Ornithorhynchus anatinus	O
CHICK\|Ensembl=ENSGALG00010028174.1\|UniProtKB=A0A8V1ACT2	Gallus gallus	O
ANOCA\|Ensembl=ENSACAG00000007523.3\|UniProtKB=G1KGR0	Anolis carolinensis	O
XENLA\|Gene=gmppb-a\|UniProtKB=Q68EY9	Xenopus laevis	O
XENLA\|Gene=gmppb-b\|UniProtKB=A2VD83	Xenopus laevis	O
XENLA\|Gene=gmppb.S\|UniProtKB=A0A1L8GHE1	Xenopus laevis	O
XENLA\|Gene=gmppb.L\|UniProtKB=A0A1L8GPA3	Xenopus laevis	O
XENTR\|Xenbase=XB-GENE-922650\|UniProtKB=Q68EQ1	Xenopus tropicalis	O
ORYLA\|Ensembl=ENSORLG00000017452.2\|UniProtKB=H2MSS8	Oryzias latipes	O
DANRE\|ZFIN=ZDB-GENE-040801-234\|UniProtKB=Q6DBU5	Danio rerio	O
LEPOC\|Ensembl=ENSLOCG00000013363.1\|UniProtKB=W5N768	lepisosteus oculatus	O
BRAFL\|Gene=LOC118416013\|UniProtKB=A0A9J7MRV8	Branchiostoma floridae	O
STRPU\|EnsemblGenome=LOC575693\|UniProtKB=A0A7M7LLH4	Strongylocentrotus purpuratus	O
CAEEL\|WormBase=WBGene00016583\|UniProtKB=A3QMC8	Caenorhabditis elegans	O
CAEBR\|WormBase=WBGene00028569\|UniProtKB=Q61S97	Caenorhabditis briggsae	O
PRIPA\|EnsemblGenome=WBGene00103230\|UniProtKB=A0A2A6BVU7	Pristionchus pacificus	O
DROME\|FlyBase=FBgn0037279\|UniProtKB=Q7JZB4	Drosophila melanogaster	O
ANOGA\|EnsemblGenome=AGAP001299\|UniProtKB=A7UVP8	Anopheles gambiae	O
TRICA\|EnsemblGenome=TC015659\|UniProtKB=D2A681	Tribolium castaneum	O
DAPPU\|EnsemblGenome=DAPPUDRAFT_101964\|UniProtKB=E9GF00	Daphnia pulex	O
IXOSC\|EnsemblGenome=ISCW019597\|UniProtKB=B7PWY1	Ixodes scapularis	O
HELRO\|EnsemblGenome=HelroG106916\|UniProtKB=T1EE59	helobdella robusta	O
NEMVE\|EnsemblGenome=NEMVEDRAFT_v1g162365\|UniProtKB=A7RT58	Nematostella vectensis	O
TRIAD\|EnsemblGenome=TriadG33563\|UniProtKB=B3SCS7	Trichoplax adhaerens	O
MONBE\|EnsemblGenome=MONBRDRAFT_12046\|UniProtKB=A9VB22	Monosiga brevicollis	O
YEAST\|SGD=S000002213\|UniProtKB=P41940	Saccharomyces cerevisiae	O
EREGS\|EnsemblGenome=AGOS_AFR599W\|UniProtKB=Q752H4	Eremothecium gossypii	O
CANAL\|CGD=CAL0000178573\|UniProtKB=O93827	Candida albicans	O
YARLI\|EnsemblGenome=YALI0_C06490g\|UniProtKB=Q6CCU3	Yarrowia lipolytica	O
EMENI\|EnsemblGenome=ANIA_05586\|UniProtKB=Q5B1J4	Emericella nidulans	O
ASPFU\|Gene_ORFName=AFUA_4G11510\|UniProtKB=Q4U3E8	Neosartorya fumigata	O
NEUCR\|EnsemblGenome=NCU06003\|UniProtKB=Q7RVR8	Neurospora crassa	O
SCLS1\|EnsemblGenome=SS1G_00164\|UniProtKB=A7E4E2	Sclerotinia sclerotiorum	O
SCHPO\|PomBase=SPCC1906.01\|UniProtKB=O74484	Schizosaccharomyces pombe	O
SCHJY\|EnsemblGenome=SJAG_03188\|UniProtKB=B6K3K0	Schizosaccharomyces japonicus	O
CRYNJ\|EnsemblGenome=CNC03020\|UniProtKB=P0CO20	Cryptococcus neoformans	O
PUCGT\|EnsemblGenome=PGTG_11653\|UniProtKB=E3KNM2	Puccinia graminis	O
BATDJ\|Gene_ORFName=BATDEDRAFT_16693\|UniProtKB=F4P2M7	Batrachochytrium dendrobatidis	O
BATDJ\|Gene_ORFName=BATDEDRAFT_20856\|UniProtKB=F4PCF5	Batrachochytrium dendrobatidis	O
DICDI\|dictyBase=DDB_G0287619\|UniProtKB=Q54K39	Dictyostelium discoideum	O
DICPU\|EnsemblGenome=DICPUDRAFT_54721\|UniProtKB=F0ZIF3	Dictyostelium purpureum	O
LEIMA\|EnsemblGenome=LMJF_23_0110\|UniProtKB=Q4QBG5	Leishmania major	O
TRYB2\|Gene_ORFName=Tb927.8.2050\|UniProtKB=Q581X7	Trypanosoma brucei	O
GIAIC\|EnsemblGenome=GL50803_16598\|UniProtKB=A8B958	Giardia intestinalis	O
TRIV3\|Gene_ORFName=TVAG_394720\|UniProtKB=A2EDD6	Trichomonas vaginalis	O
TRIV3\|Gene_ORFName=TVAG_397470\|UniProtKB=A2G1C4	Trichomonas vaginalis	O
TRIV3\|Gene_ORFName=TVAG_178210\|UniProtKB=A2DIH3	Trichomonas vaginalis	O
PLAF7\|EnsemblGenome=PF3D7_1420900\|UniProtKB=Q8ILP1	Plasmodium falciparum	O
PARTE\|EnsemblGenome=GSPATT00002821001\|UniProtKB=A0DQV4	Paramecium tetraurelia	O
PARTE\|EnsemblGenome=GSPATT00001206001\|UniProtKB=A0CW37	Paramecium tetraurelia	O
PHYRM\|Gene=H3GA81_PHYRM\|UniProtKB=H3GA81	Phytophthora ramorum	O
CHLRE\|EnsemblGenome=CHLRE_16g672800v5\|UniProtKB=A8J3F9	Chlamydomonas reinhardtii	O
AMBTC\|EnsemblGenome=AMTR_s00012p00262630\|UniProtKB=W1PDN1	Amborella trichopoda	O
AMBTC\|EnsemblGenome=AMTR_s00012p00262290\|UniProtKB=W1PJY2	Amborella trichopoda	O
ZOSMR\|Gene_ORFName=ZOSMA_56G00970\|UniProtKB=A0A0K9NVS2	Zostera marina	O
ZOSMR\|Gene_ORFName=ZOSMA_93G00280\|UniProtKB=A0A0K9NII3	Zostera marina	O
MUSAM\|EnsemblGenome=Ma02_g21000\|UniProtKB=A0A804I550	Musa acuminata subsp. malaccensis	O
SETIT\|EnsemblGenome=SETIT_036322mg\|UniProtKB=K4ABS0	Setaria italica	O
SORBI\|EnsemblGenome=SORBI_3001G424500\|UniProtKB=A0A1B6QP69	Sorghum bicolor	O
MAIZE\|EnsemblGenome=Zm00001eb012020\|UniProtKB=B4G0L2	Zea mays	O
ORYSJ\|EnsemblGenome=Os03g0268400\|UniProtKB=Q84JH5	Oryza sativa	O
BRADI\|EnsemblGenome=BRADI_1g66930v3\|UniProtKB=I1H726	Brachypodium distachyon	O
HORVV\|EnsemblGenome=HORVU.MOREX.r3.4HG0389060\|UniProtKB=M0YL77	Hordeum vulgare subsp. vulgare	O
WHEAT\|EnsemblGenome=TraesCS4A02G094000\|UniProtKB=A0A096UQP3	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS4B02G210300\|UniProtKB=A0A3B6IQV6	Triticum aestivum	O
MUSAM\|EnsemblGenome=Ma02_g18300\|UniProtKB=A0A804I447	Musa acuminata subsp. malaccensis	O
MUSAM\|EnsemblGenome=Ma08_g02320\|UniProtKB=A0A804K226	Musa acuminata subsp. malaccensis	O
MUSAM\|EnsemblGenome=Ma06_g35930\|UniProtKB=A0A804JP78	Musa acuminata subsp. malaccensis	O
SETIT\|EnsemblGenome=SETIT_010432mg\|UniProtKB=K3Y891	Setaria italica	O
SETIT\|EnsemblGenome=SETIT_001333mg\|UniProtKB=K3XHG2	Setaria italica	O
SORBI\|EnsemblGenome=SORBI_3003G355200\|UniProtKB=C5XPS1	Sorghum bicolor	O
MAIZE\|EnsemblGenome=Zm00001eb367780\|UniProtKB=A0A804QYM9	Zea mays	O
MAIZE\|EnsemblGenome=Zm00001eb148030\|UniProtKB=C4JAC1	Zea mays	O
ORYSJ\|EnsemblGenome=Os01g0847200\|UniProtKB=Q941T9	Oryza sativa	O
ORYSJ\|EnsemblGenome=Os08g0237200\|UniProtKB=Q6Z9A3	Oryza sativa	O
BRADI\|EnsemblGenome=BRADI_2g54870v3\|UniProtKB=I1HTC9	Brachypodium distachyon	O
HORVV\|EnsemblGenome=HORVU.MOREX.r3.3HG0302100\|UniProtKB=F2D8Y2	Hordeum vulgare subsp. vulgare	O
WHEAT\|EnsemblGenome=TraesCS3B02G395400\|UniProtKB=W5CPQ7	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS3D02G356700\|UniProtKB=A0A3B6H1S9	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS3A02G363000\|UniProtKB=A0A3B6EQD2	Triticum aestivum	O
NELNU\|Gene_ORFName=HUJ06_029777\|UniProtKB=A0A1U8AK20	Nelumbo nucifera	O
NELNU\|Gene_ORFName=HUJ06_019610\|UniProtKB=A0A1U8B3B5	Nelumbo nucifera	O
NELNU\|Gene=LOC104597092\|UniProtKB=A0A1U7ZTA6	Nelumbo nucifera	O
SPIOL\|Gene=LOC110785479\|UniProtKB=A0A9R0ICI5	Spinacia oleracea	O
HELAN\|EnsemblGenome=HanXRQr2_Chr14g0665091\|UniProtKB=A0A251TQ17	Helianthus annuus	O
HELAN\|Gene_ORFName=HannXRQ_Chr07g0201721\|UniProtKB=A0A251UDV0	Helianthus annuus	O
HELAN\|Gene_ORFName=HanXRQr2_Chr07g0311221\|UniProtKB=A0A9K3NGV9	Helianthus annuus	O
LACSA\|Gene_ORFName=LSAT_V11C200093750\|UniProtKB=A0A9R1WJV6	Lactuca sativa	O
HELAN\|Gene_ORFName=HannXRQ_Chr09g0247091\|UniProtKB=A0A251TUI4	Helianthus annuus	O
HELAN\|Gene_ORFName=HanXRQr2_Chr09g0381671\|UniProtKB=A0A9K3N800	Helianthus annuus	O
HELAN\|Gene_ORFName=HanXRQr2_Chr15g0721161\|UniProtKB=A0A9K3E5T0	Helianthus annuus	O
HELAN\|Gene_ORFName=HannXRQ_Chr15g0496461\|UniProtKB=A0A251SCR9	Helianthus annuus	O
LACSA\|Gene_ORFName=LSAT_V11C900482670\|UniProtKB=A0A9R1UIZ5	Lactuca sativa	O
LACSA\|Gene_ORFName=LSAT_V11C200093750\|UniProtKB=A0A9R1WJV6	Lactuca sativa	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a008833mg\|UniProtKB=A0A022PUB6	Erythranthe guttata	O
CAPAN\|EnsemblGenome=T459_08997\|UniProtKB=A0A2G2ZY63	Capsicum annuum	O
SOLTU\|EnsemblGenome=PGSC0003DMG400015098\|UniProtKB=Q9ZTW5	Solanum tuberosum	O
SOLLC\|EnsemblGenome=Solyc03g096730.3\|UniProtKB=Q6J1L7	Solanum lycopersicum	O
TOBAC\|Gene=LOC107788373\|UniProtKB=A0A1S3ZMQ1	Nicotiana tabacum	O
TOBAC\|Gene=LOC107810135\|UniProtKB=A0A1S4BNB5	Nicotiana tabacum	O
SOLTU\|EnsemblGenome=PGSC0003DMG400015645\|UniProtKB=M1B9V2	Solanum tuberosum	O
SOLLC\|EnsemblGenome=Solyc06g051270.3\|UniProtKB=A0A3Q7GS00	Solanum lycopersicum	O
TOBAC\|Gene=LOC107771893\|UniProtKB=A0A1S3Y3W2	Nicotiana tabacum	O
TOBAC\|Gene=LOC107770019\|UniProtKB=A0A1S3XXZ8	Nicotiana tabacum	O
EUCGR\|EnsemblGenome=EUGRSUZ_G02988\|UniProtKB=A0A059BI79	Eucalyptus grandis	O
BRANA\|Gene_ORFName=DARMORV10_C04P07730.1\|UniProtKB=A0A078G1Q2	Brassica napus	O
BRACM\|EnsemblGenome=Bra005014\|UniProtKB=M4CLC7	Brassica campestris	O
BRANA\|Gene_ORFName=GSBRNA2T00124741001\|UniProtKB=A0A078F2P1	Brassica napus	O
BRANA\|Gene_ORFName=GSBRNA2T00034358001\|UniProtKB=A0A078JBH7	Brassica napus	O
BRACM\|EnsemblGenome=Bra017052\|UniProtKB=A0A397ZNX0	Brassica campestris	O
BRACM\|EnsemblGenome=Bra000149\|UniProtKB=M4C7H1	Brassica campestris	O
ARATH\|TAIR=locus=2100001\|UniProtKB=Q9M2S0	Arabidopsis thaliana	O
ARATH\|TAIR=locus=2005504\|UniProtKB=O22287	Arabidopsis thaliana	O
THECC\|EnsemblGenome=TCM_042692\|UniProtKB=A0A061FL22	Theobroma cacao	O
CITSI\|Gene_ORFName=CISIN_1g018060mg\|UniProtKB=A0A067FP88	Citrus sinensis	O
POPTR\|EnsemblGenome=POPTR_008G060100v3\|UniProtKB=B9HMM2	Populus trichocarpa	O
POPTR\|EnsemblGenome=POPTR_008G006700v3\|UniProtKB=B9HK47	Populus trichocarpa	O
POPTR\|EnsemblGenome=POPTR_010G198800v3\|UniProtKB=B9HT64	Populus trichocarpa	O
MANES\|EnsemblGenome=MANES_07G120300\|UniProtKB=A0A2C9VN57	Manihot esculenta	O
MANES\|EnsemblGenome=MANES_10G025000\|UniProtKB=A0A2C9V484	Manihot esculenta	O
RICCO\|Gene_ORFName=RCOM_1934400\|UniProtKB=B9TF87	Ricinus communis	O
RICCO\|Gene_ORFName=RCOM_1482320\|UniProtKB=B9SE08	Ricinus communis	O
MEDTR\|EnsemblGenome=MTR_3g069070\|UniProtKB=G7J9Y8	Medicago truncatula	O
MEDTR\|EnsemblGenome=MTR_4g133690\|UniProtKB=G7JLR8	Medicago truncatula	O
MEDTR\|EnsemblGenome=MTR_5g080770\|UniProtKB=G7KGV6	Medicago truncatula	O
JUGRE\|EnsemblGenome=Jr12_23660\|UniProtKB=A0A2I4HMU2	Juglans regia	O
CUCSA\|EnsemblGenome=Csa_6G008750\|UniProtKB=A0A0A0KA78	Cucumis sativus	O
PRUPE\|EnsemblGenome=PRUPE_2G240500\|UniProtKB=B6ZL91	Prunus persica	O
SELML\|EnsemblGenome=SELMODRAFT_185648\|UniProtKB=D8T5L0	Selaginella moellendorffii	O
SELML\|EnsemblGenome=SELMODRAFT_267755\|UniProtKB=D8RTK5	Selaginella moellendorffii	O
SELML\|EnsemblGenome=SELMODRAFT_168373\|UniProtKB=D8R6R6	Selaginella moellendorffii	O
PHYPA\|EnsemblGenome=Pp3c24_16870\|UniProtKB=A9TU90	Physcomitrella patens	O
PHYPA\|EnsemblGenome=Pp3c8_17380\|UniProtKB=A9SSX9	Physcomitrella patens	O
MARPO\|EnsemblGenome=MARPO_0034s0043\|UniProtKB=A0A1L7B546	Marchantia polymorpha	O
KLENI\|Gene_ORFName=KFL_008460020\|UniProtKB=A0A1Y1IM91	Klebsormidium nitens	O
NITMS\|EnsemblGenome=Nmar_0424\|UniProtKB=A9A5C1	Nitrosopumilus maritimus	O
NITMS\|EnsemblGenome=Nmar_0843\|UniProtKB=A9A5M8	Nitrosopumilus maritimus	O
SACS2\|EnsemblGenome=SSO0317\|UniProtKB=Q980H9	Sulfolobus solfataricus	O
PYRAE\|EnsemblGenome=PAE3554\|UniProtKB=Q8ZSW5	Pyrobaculum aerophilum	O
SACS2\|EnsemblGenome=SSO0381\|UniProtKB=Q980D4	Sulfolobus solfataricus	O
SACS2\|EnsemblGenome=SSO2479\|UniProtKB=Q97VX4	Sulfolobus solfataricus	O
HALSA\|EnsemblGenome=VNG_0390G\|UniProtKB=Q9HS61	Halobacterium salinarum	O
THEKO\|EnsemblGenome=TK1711\|UniProtKB=Q5JJ02	Thermococcus kodakaraensis	O
METAC\|EnsemblGenome=MA_3140\|UniProtKB=Q8TL99	Methanosarcina acetivorans	O
THEKO\|EnsemblGenome=TK0955\|UniProtKB=Q5JI97	Thermococcus kodakaraensis	O
CHLAA\|EnsemblGenome=Caur_2687\|UniProtKB=A9WJI8	Chloroflexus aurantiacus	O
STRCO\|Gene_OrderedLocusName=SCO3039\|UniProtKB=Q9KZK6	Streptomyces coelicolor	O
MYCTU\|Gene_OrderedLocusName=Rv3264c\|UniProtKB=L7N6A5	Mycobacterium tuberculosis	O
CHLAA\|EnsemblGenome=Caur_1473\|UniProtKB=A9WAE2	Chloroflexus aurantiacus	O
CHLAA\|EnsemblGenome=Caur_1759\|UniProtKB=A9WCI5	Chloroflexus aurantiacus	O
CHLAA\|EnsemblGenome=Caur_2960\|UniProtKB=A9WG05	Chloroflexus aurantiacus	O
STRCO\|Gene_OrderedLocusName=SCO1388\|UniProtKB=Q9KZP4	Streptomyces coelicolor	O
BACCR\|EnsemblGenome=BC_4264\|UniProtKB=Q818I9	Bacillus cereus	O
GLOVI\|Gene=Q7NII3_GLOVI\|UniProtKB=Q7NII3	Gloeobacter violaceus	O
GLOVI\|Gene=Q7NF24_GLOVI\|UniProtKB=Q7NF24	Gloeobacter violaceus	O
GLOVI\|Gene=rfbM\|UniProtKB=Q7NJQ0	Gloeobacter violaceus	O
GLOVI\|Gene=Q7NNE0_GLOVI\|UniProtKB=Q7NNE0	Gloeobacter violaceus	O
GLOVI\|Gene=Q7NC95_GLOVI\|UniProtKB=Q7NC95	Gloeobacter violaceus	O
SYNY3\|EnsemblGenome=BAA18379\|UniProtKB=P74285	Synechocystis	O
GLOVI\|Gene=Q7NG32_GLOVI\|UniProtKB=Q7NG32	Gloeobacter violaceus	O
SYNY3\|Gene=P74589_SYNY3\|UniProtKB=P74589	Synechocystis	O
LEPIN\|EnsemblGenome=LA_1581\|UniProtKB=Q8F5T6	Leptospira interrogans	O
LEPIN\|EnsemblGenome=LA_1611\|UniProtKB=Q8F5Q6	Leptospira interrogans	O
AQUAE\|EnsemblGenome=aq_718\|UniProtKB=O66933	Aquifex aeolicus	O
DICTD\|EnsemblGenome=Dtur_1427\|UniProtKB=B8E0W4	Dictyoglomus turgidum	O
GEOSL\|EnsemblGenome=GSU1968\|UniProtKB=Q74B34	Geobacter sulfurreducens	O
GEOSL\|EnsemblGenome=GSU3254\|UniProtKB=Q747L1	Geobacter sulfurreducens	O
BRADU\|Gene=Q89HJ6_BRADU\|UniProtKB=Q89HJ6	Bradyrhizobium diazoefficiens	O