PANTHER Gene Information

Gene Symbol(s):

TRIADDRAFT_53152

Organism:

Trichoplax adhaerens

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Tree

Gene Name:

Adenylate kinase

Gene ID:

TriadG53152

Protein ID:

B3RNF9

Persistent Id:

PTN001156918

Alternate Ids:

6751064(GeneID)	195998824(GI)
TriadT53152(Ensembl_TRS)	TriadP53152(Ensembl_PRO)
EDV27446(EMBL-CDS)	B3RNF9(Synonym)
TriadG53152(Ensembl)	DS985242(EMBL)
B3RNF9_TRIAD(UniProtKB-ID)	XP_002109280(refSeq)
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PANTHER Classification

PANTHER Family:	NUCLEOTIDE KINASE (PTHR23359)
PANTHER Subfamily:	ADENYLATE KINASE 8 (PTHR23359:SF167)
PANTHER GO-slim Molecular Function:	nucleoside diphosphate kinase activity
PANTHER GO-slim Biological Process:
PANTHER GO-slim Cellular Component:	cytoplasm
PANTHER protein class:	nucleotide kinase

Pathway Categories:

De novo purine biosynthesis

Adenylate kinase

Salvage pyrimidine ribonucleotides

Cytidylate kinase

GENE ONTOLOGY DATABASE ANNOTATIONS

Click to Expand...

GO MF Complete:	adenylate kinase activity, ATP binding, kinase activity
GO BP Complete:	nucleoside monophosphate phosphorylation, nucleobase-containing compound metabolic process
GO CC Complete:	cytoplasm, cytosol

REACTOME DATABASE ANNOTATIONS Click to Expand...

Reactome Pathways:

Orthologs

ID	Organism	Type
HUMAN\|HGNC=26526\|UniProtKB=Q96MA6	Homo sapiens	LDO
PANTR\|Ensembl=ENSPTRG00000021502.4\|UniProtKB=H2QY34	Pan troglodytes	LDO
GORGO\|Ensembl=ENSGGOG00000002135.3\|UniProtKB=G3QI98	Gorilla gorilla gorilla	LDO
MACMU\|Ensembl=ENSMMUG00000009916.4\|UniProtKB=A0A5F8A4H7	Macaca mulatta	LDO
MOUSE\|MGI=MGI=1916120\|UniProtKB=Q32M07	Mus musculus	LDO
RAT\|RGD=1303144\|UniProtKB=Q68FP8	Rattus norvegicus	LDO
BOVIN\|Ensembl=ENSBTAG00000004092.5\|UniProtKB=E1BLG0	Bos taurus	LDO
PIG\|Ensembl=ENSSSCG00015061468\|UniProtKB=A0A4X1T4U3	Sus scrofa	LDO
PIG\|Ensembl=ENSSSCG00000039188.3\|UniProtKB=A0A287AID3	Sus scrofa	LDO
HORSE\|Ensembl=ENSECAG00000006330.4\|UniProtKB=A0A3Q2GUX0	Equus caballus	LDO
FELCA\|Ensembl=ENSFCAG00000023153.4\|UniProtKB=A0A2I2V3K2	Felis catus	LDO
CANLF\|Ensembl=ENSCAFG00845016359.1\|UniProtKB=A0A8I3NQB2	Canis lupus familiaris	LDO
MONDO\|Ensembl=ENSMODG00000012561.4\|UniProtKB=F6PLL4	Monodelphis domestica	LDO
ORNAN\|Ensembl=ENSOANG00000005052.3\|UniProtKB=F6Q864	Ornithorhynchus anatinus	LDO
CHICK\|Ensembl=ENSGALG00010028800.1\|UniProtKB=A0A8V1AJZ1	Gallus gallus	LDO
XENLA\|Gene=ak8.L\|UniProtKB=A0A8J1LG97	Xenopus laevis	LDO
XENLA\|Gene=ak8\|UniProtKB=Q5M7G4	Xenopus laevis	LDO
XENTR\|Xenbase=XB-GENE-5831327\|UniProtKB=Q6P618	Xenopus tropicalis	LDO
ORYLA\|Ensembl=ENSORLG00000016621.2\|UniProtKB=H2MPY9	Oryzias latipes	LDO
DANRE\|ZFIN=ZDB-GENE-050522-275\|UniProtKB=Q502L7	Danio rerio	LDO
LEPOC\|Ensembl=ENSLOCG00000005659.1\|UniProtKB=W5MEM4	lepisosteus oculatus	LDO
CIOIN\|Ensembl=ENSCING00000004295.3\|UniProtKB=C4B8W9	Ciona intestinalis	LDO
BRAFL\|Gene=LOC118420470\|UniProtKB=A0A9J7LI48	Branchiostoma floridae	LDO
STRPU\|EnsemblGenome=LOC584538\|UniProtKB=A0A7M7P7G5	Strongylocentrotus purpuratus	LDO
DROME\|FlyBase=FBgn0039799\|UniProtKB=Q9VA43	Drosophila melanogaster	LDO
NEMVE\|EnsemblGenome=NEMVEDRAFT_v1g188080\|UniProtKB=A7SC73	Nematostella vectensis	O
NEMVE\|EnsemblGenome=NEMVEDRAFT_v1g98446\|UniProtKB=A7RYZ3	Nematostella vectensis	LDO
MONBE\|EnsemblGenome=MONBRDRAFT_30971\|UniProtKB=A9UQG8	Monosiga brevicollis	LDO
LEIMA\|EnsemblGenome=LMJF_36_1360\|UniProtKB=Q4Q1U0	Leishmania major	LDO
TRYB2\|Gene_ORFName=Tb10.70.7330\|UniProtKB=Q38CF7	Trypanosoma brucei	O
TRYB2\|Gene_ORFName=Tb10.70.1200\|UniProtKB=Q38B28	Trypanosoma brucei	LDO
TRIV3\|Gene_ORFName=TVAG_282000\|UniProtKB=A2E9R5	Trichomonas vaginalis	O
TRIV3\|Gene_ORFName=TVAG_372350\|UniProtKB=A2G001	Trichomonas vaginalis	LDO
PHYRM\|Gene=H3GGY7_PHYRM\|UniProtKB=H3GGY7	Phytophthora ramorum	LDO
CHLRE\|EnsemblGenome=CHLRE_07g325723v5\|UniProtKB=A8JH21	Chlamydomonas reinhardtii	LDO
AMBTC\|EnsemblGenome=AMTR_s00026p00112660\|UniProtKB=W1PK61	Amborella trichopoda	LDO
ZOSMR\|Gene_ORFName=ZOSMA_96G00320\|UniProtKB=A0A0K9NIA7	Zostera marina	LDO
SETIT\|EnsemblGenome=SETIT_013366mg\|UniProtKB=K3YGJ7	Setaria italica	LDO
SORBI\|EnsemblGenome=SORBI_3007G113600\|UniProtKB=A0A1B6PH86	Sorghum bicolor	LDO
MAIZE\|EnsemblGenome=Zm00001eb177670\|UniProtKB=C0P8P8	Zea mays	LDO
ORYSJ\|Gene_OrderedLocusName=LOC_Os08g19140\|UniProtKB=Q0J6P7	Oryza sativa	LDO
BRADI\|EnsemblGenome=BRADI_3g19850v3\|UniProtKB=I1I2L8	Brachypodium distachyon	LDO
HORVV\|EnsemblGenome=HORVU.MOREX.r3.6HG0551710\|UniProtKB=F2D6P8	Hordeum vulgare subsp. vulgare	LDO
WHEAT\|EnsemblGenome=TraesCS6B02G113200\|UniProtKB=A0A3B6PHE7	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS6A02G083900\|UniProtKB=A0A3B6NKU3	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS6D02G077500\|UniProtKB=A0A3B6QDA8	Triticum aestivum	LDO
NELNU\|Gene_ORFName=HUJ06_003370\|UniProtKB=A0A1U7ZBZ0	Nelumbo nucifera	LDO
SPIOL\|Gene=LOC110790117\|UniProtKB=A0A9R0JXC1	Spinacia oleracea	LDO
HELAN\|Gene_ORFName=HanXRQr2_Chr10g0461311\|UniProtKB=A0A9K3I059	Helianthus annuus	LDO
LACSA\|Gene_ORFName=LSAT_V11C500261780\|UniProtKB=A0A9R1VKW7	Lactuca sativa	LDO
ERYGU\|Gene_ORFName=MIMGU_mgv1a0039952mg\|UniProtKB=A0A022QZ26	Erythranthe guttata	LDO
CAPAN\|EnsemblGenome=T459_24499\|UniProtKB=A0A2G2YVI2	Capsicum annuum	LDO
SOLLC\|EnsemblGenome=Solyc12g010380.2\|UniProtKB=A0A3Q7J418	Solanum lycopersicum	LDO
TOBAC\|Gene=LOC107770764\|UniProtKB=A0A1S3Y0E5	Nicotiana tabacum	O
TOBAC\|Gene=LOC107778794\|UniProtKB=A0A1S3YQU6	Nicotiana tabacum	LDO
VITVI\|EnsemblGenome=Vitvi11g04151\|UniProtKB=F6HCJ8	Vitis vinifera	LDO
EUCGR\|EnsemblGenome=EUGRSUZ_C00416\|UniProtKB=A0A059CKK7	Eucalyptus grandis	LDO
BRANA\|Gene_ORFName=GSBRNA2T00023626001\|UniProtKB=A0A078J3S0	Brassica napus	O
BRANA\|Gene_ORFName=GSBRNA2T00073632001\|UniProtKB=A0A078HT82	Brassica napus	LDO
BRACM\|EnsemblGenome=Bra038939\|UniProtKB=M4FCW9	Brassica campestris	LDO
ARATH\|TAIR=locus=2182407\|UniProtKB=Q8VYL1	Arabidopsis thaliana	LDO
GOSHI\|Gene=LOC107901372\|UniProtKB=A0A1U8J3P4	Gossypium hirsutum	O
GOSHI\|Gene=LOC107962538\|UniProtKB=A0A1U8PVQ7	Gossypium hirsutum	LDO
THECC\|EnsemblGenome=TCM_038279\|UniProtKB=A0A061GW31	Theobroma cacao	LDO
POPTR\|EnsemblGenome=POPTR_018G113400v3\|UniProtKB=A0A2K1WZG8	Populus trichocarpa	LDO
MANES\|EnsemblGenome=MANES_14G139700\|UniProtKB=A0A2C9ULQ3	Manihot esculenta	LDO
RICCO\|Gene_ORFName=RCOM_0701690\|UniProtKB=B9SC22	Ricinus communis	LDO
SOYBN\|EnsemblGenome=GLYMA_09G096800\|UniProtKB=K7LCU0	Glycine max	O
SOYBN\|EnsemblGenome=GLYMA_15G203100\|UniProtKB=I1MI09	Glycine max	LDO
MEDTR\|EnsemblGenome=MTR_2g461290\|UniProtKB=A0A072V8Z3	Medicago truncatula	LDO
JUGRE\|EnsemblGenome=Jr15_13310\|UniProtKB=A0A2I4FQ57	Juglans regia	LDO
CUCSA\|EnsemblGenome=Csa_2G008770\|UniProtKB=A0A0A0LJ30	Cucumis sativus	LDO
PRUPE\|EnsemblGenome=PRUPE_7G253500\|UniProtKB=A0A251NGT6	Prunus persica	LDO
LEIMA\|EnsemblGenome=LMJF_36_6030\|UniProtKB=Q4Q0F7	Leishmania major	O
TRYB2\|Gene_ORFName=Tb927.5.3970\|UniProtKB=Q57U69	Trypanosoma brucei	O
CHLRE\|EnsemblGenome=CHLRE_06g249900v5\|UniProtKB=A8HYW3	Chlamydomonas reinhardtii	O
CAPAN\|EnsemblGenome=T459_34454\|UniProtKB=A0A2G2XWK4	Capsicum annuum	O
SOLTU\|EnsemblGenome=PGSC0003DMG400026110\|UniProtKB=M1CGT0	Solanum tuberosum	O
SOLLC\|EnsemblGenome=Solyc06g065270.3\|UniProtKB=A0A3Q7GWK3	Solanum lycopersicum	O
SELML\|EnsemblGenome=SELMODRAFT_169443\|UniProtKB=D8R8W9	Selaginella moellendorffii	O
SELML\|EnsemblGenome=SELMODRAFT_270406\|UniProtKB=D8QYR9	Selaginella moellendorffii	O
LEIMA\|EnsemblGenome=LMJF_21_1250\|UniProtKB=Q4QC71	Leishmania major	O
TRYB2\|Gene_ORFName=Tb927.2.5660\|UniProtKB=Q586D5	Trypanosoma brucei	O
PARTE\|EnsemblGenome=GSPATT00007605001\|UniProtKB=A0CH98	Paramecium tetraurelia	O
PARTE\|EnsemblGenome=GSPATT00014845001\|UniProtKB=A0DA71	Paramecium tetraurelia	O
PARTE\|EnsemblGenome=GSPATT00038139001\|UniProtKB=A0CFY0	Paramecium tetraurelia	O
THAPS\|EnsemblGenome=THAPSDRAFT_9511\|UniProtKB=B8CBJ0	Thalassiosira pseudonana	O
PHYRM\|Gene=H3GIP3_PHYRM\|UniProtKB=H3GIP3	Phytophthora ramorum	O
PHYRM\|Gene=H3GIP9_PHYRM\|UniProtKB=H3GIP9	Phytophthora ramorum	O
CHLRE\|EnsemblGenome=CHLRE_12g494850v5\|UniProtKB=A8JDN2	Chlamydomonas reinhardtii	O
AMBTC\|EnsemblGenome=AMTR_s00071p00171040\|UniProtKB=U5DC05	Amborella trichopoda	O
ZOSMR\|Gene_ORFName=ZOSMA_161G00420\|UniProtKB=A0A0K9PWG5	Zostera marina	O
ZOSMR\|Gene_ORFName=ZOSMA_105G00760\|UniProtKB=A0A0K9Q668	Zostera marina	O
MUSAM\|EnsemblGenome=Ma02_g02660\|UniProtKB=A0A804HYK4	Musa acuminata subsp. malaccensis	O
MUSAM\|EnsemblGenome=Ma07_g24940\|UniProtKB=A0A804JZJ5	Musa acuminata subsp. malaccensis	O
SETIT\|EnsemblGenome=SETIT_014186mg\|UniProtKB=K3YIW7	Setaria italica	O
SORBI\|EnsemblGenome=SORBI_3007G009200\|UniProtKB=C5YLV3	Sorghum bicolor	O
MAIZE\|EnsemblGenome=Zm00001eb260410\|UniProtKB=B4FL09	Zea mays	O
MAIZE\|Gene=ADK1\|UniProtKB=P43188	Zea mays	O
ORYSJ\|EnsemblGenome=Os08g0109300\|UniProtKB=Q6ZC69	Oryza sativa	O
BRADI\|EnsemblGenome=BRADI_3g13360v3\|UniProtKB=I1I0D2	Brachypodium distachyon	O
HORVV\|EnsemblGenome=HORVU.MOREX.r3.7HG0706290\|UniProtKB=A0A8I6YG19	Hordeum vulgare subsp. vulgare	O
WHEAT\|EnsemblGenome=TraesCS7D02G319000\|UniProtKB=A0A3B6TSL3	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS7B02G223000\|UniProtKB=A0A3B6SKW7	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS7A02G322100\|UniProtKB=A0A3B6RKM9	Triticum aestivum	O
NELNU\|Gene=LOC104586485\|UniProtKB=A0A1U7Z407	Nelumbo nucifera	O
SPIOL\|Gene=LOC110790577\|UniProtKB=A0A9R0JXS9	Spinacia oleracea	O
SPIOL\|Gene=LOC110786006\|UniProtKB=A0A9R0IBA7	Spinacia oleracea	O
HELAN\|EnsemblGenome=HanXRQr2_Chr02g0064491\|UniProtKB=A0A251VEG4	Helianthus annuus	O
LACSA\|Gene_ORFName=LSAT_V11C100004670\|UniProtKB=A0A9R1WSV0	Lactuca sativa	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a013591mg\|UniProtKB=A0A022Q6G9	Erythranthe guttata	O
CAPAN\|EnsemblGenome=T459_30832\|UniProtKB=A0A2G2Y9H2	Capsicum annuum	O
CAPAN\|EnsemblGenome=T459_01204\|UniProtKB=A0A2G3AGF2	Capsicum annuum	O
SOLTU\|EnsemblGenome=PGSC0003DMG400013933\|UniProtKB=M1B3D6	Solanum tuberosum	O
TOBAC\|Gene=LOC107784862\|UniProtKB=A0A1S3ZBD8	Nicotiana tabacum	O
TOBAC\|Gene=LOC107805684\|UniProtKB=A0A1S4B8X3	Nicotiana tabacum	O
VITVI\|EnsemblGenome=Vitvi18g04574\|UniProtKB=F6I3V3	Vitis vinifera	O
EUCGR\|EnsemblGenome=EUGRSUZ_F02985\|UniProtKB=A0A059BTH2	Eucalyptus grandis	O
BRANA\|Gene_ORFName=DARMORV10_A09P04250.1\|UniProtKB=A0A078G4T3	Brassica napus	O
BRANA\|Gene_ORFName=GSBRNA2T00038056001\|UniProtKB=A0A078JB69	Brassica napus	O
BRACM\|EnsemblGenome=Bra037526\|UniProtKB=M4F8W4	Brassica campestris	O
BRACM\|EnsemblGenome=Bra036193\|UniProtKB=M4F537	Brassica campestris	O
ARATH\|TAIR=locus=2160942\|UniProtKB=Q9FIJ7	Arabidopsis thaliana	O
GOSHI\|Gene=LOC107957472\|UniProtKB=A0A1U8PBA7	Gossypium hirsutum	O
GOSHI\|Gene=LOC107904874\|UniProtKB=A0A1U8J976	Gossypium hirsutum	O
THECC\|EnsemblGenome=TCM_036205\|UniProtKB=A0A061FK41	Theobroma cacao	O
CITSI\|Gene_ORFName=CISIN_1g022042mg\|UniProtKB=A0A067DSM3	Citrus sinensis	O
POPTR\|Gene_ORFName=POPTR_019G078200\|UniProtKB=A0A3N7G8S2	Populus trichocarpa	O
POPTR\|EnsemblGenome=POPTR_013G103400v3\|UniProtKB=A0A2K1Y3T5	Populus trichocarpa	O
MANES\|EnsemblGenome=MANES_02G199500\|UniProtKB=A0A2C9WH08	Manihot esculenta	O
RICCO\|Gene_ORFName=RCOM_1330260\|UniProtKB=B9S706	Ricinus communis	O
SOYBN\|EnsemblGenome=GLYMA_09G234300\|UniProtKB=C6THI7	Glycine max	O
MEDTR\|EnsemblGenome=MTR_4g035170\|UniProtKB=G7JRB1	Medicago truncatula	O
JUGRE\|EnsemblGenome=Jr11_30750\|UniProtKB=A0A2I4E9V0	Juglans regia	O
CUCSA\|EnsemblGenome=Csa_6G052070\|UniProtKB=A0A0A0KE97	Cucumis sativus	O
PRUPE\|EnsemblGenome=PRUPE_1G556600\|UniProtKB=A0A251RIM1	Prunus persica	O
SELML\|EnsemblGenome=SELMODRAFT_77790\|UniProtKB=D8QT58	Selaginella moellendorffii	O
SELML\|EnsemblGenome=SELMODRAFT_83077\|UniProtKB=D8R2E1	Selaginella moellendorffii	O
PHYPA\|EnsemblGenome=Pp3c27_6280\|UniProtKB=A0A2K1IAX1	Physcomitrella patens	O
PHYPA\|EnsemblGenome=Pp3c6_20610\|UniProtKB=A0A7I4DZM8	Physcomitrella patens	O
MARPO\|EnsemblGenome=MARPO_0139s0019\|UniProtKB=A0A2R6W6R1	Marchantia polymorpha	O
KLENI\|Gene_ORFName=KFL_000510230\|UniProtKB=A0A1Y1HR93	Klebsormidium nitens	O
KORCO\|EnsemblGenome=Kcr_1559\|UniProtKB=B1L775	Korarchaeum cryptofilum	O
HALSA\|EnsemblGenome=VNG_1724G\|UniProtKB=Q9HPA7	Halobacterium salinarum	O
THEKO\|EnsemblGenome=TK0812\|UniProtKB=Q5JH68	Thermococcus kodakaraensis	O
CHLAA\|EnsemblGenome=Caur_2957\|UniProtKB=A9WG02	Chloroflexus aurantiacus	O
CHLAA\|EnsemblGenome=Caur_2390\|UniProtKB=A9WH87	Chloroflexus aurantiacus	O
STRCO\|Gene_OrderedLocusName=SCO4723\|UniProtKB=P43414	Streptomyces coelicolor	O
MYCTU\|Gene_OrderedLocusName=Rv0733\|UniProtKB=P9WKF5	Mycobacterium tuberculosis	O
CLOBH\|Gene_OrderedLocusName=CBO3460\|UniProtKB=A5I7I5	Clostridium botulinum	O
STRR6\|EnsemblGenome=spr0210\|UniProtKB=Q8DRD4	Streptococcus pneumoniae	O
STAA8\|EnsemblGenome=SAOUHSC_02490\|UniProtKB=Q2FW27	Staphylococcus aureus	O
LISMO\|EnsemblGenome=CAD00689\|UniProtKB=Q8Y449	Listeria monocytogenes	O
BACSU\|EnsemblGenome=BSU_01370\|UniProtKB=P16304	Bacillus subtilis	O
BACCR\|EnsemblGenome=BC_0152\|UniProtKB=Q81J22	Bacillus cereus	O
THEMA\|EnsemblGenome=TM_1479\|UniProtKB=Q9X1I8	Thermotoga maritima	O
GLOVI\|EnsemblGenome=BAC89333\|UniProtKB=Q7NKT5	Gloeobacter violaceus	O
SYNY3\|EnsemblGenome=BAA16643\|UniProtKB=P72641	Synechocystis	O
SYNY3\|EnsemblGenome=BAA17330\|UniProtKB=P73302	Synechocystis	O
LEPIN\|EnsemblGenome=LA_0760\|UniProtKB=Q9XD15	Leptospira interrogans	O
BACTN\|EnsemblGenome=BT_4387\|UniProtKB=Q89ZJ0	Bacteroides thetaiotaomicron	O
CHLTR\|EnsemblGenome=CT_128\|UniProtKB=O84130	Chlamydia trachomatis	O
RHOBA\|EnsemblGenome=RB7867\|UniProtKB=Q7UN00	Rhodopirellula baltica	O
AQUAE\|EnsemblGenome=aq_078\|UniProtKB=O66490	Aquifex aeolicus	O
THEYD\|EnsemblGenome=THEYE_A1836\|UniProtKB=B5YHP1	Thermodesulfovibrio yellowstonii	O
DICTD\|EnsemblGenome=Dtur_1001\|UniProtKB=B8E1F3	Dictyoglomus turgidum	O
FUSNN\|EnsemblGenome=FN1298\|UniProtKB=Q8RE31	Fusobacterium nucleatum	O
HELPY\|EnsemblGenome=HP_0618\|UniProtKB=P56104	Helicobacter pylori	O
NEIMB\|EnsemblGenome=NMB0823\|UniProtKB=P0A0U7	Neisseria meningitidis serogroup b	O
GEOSL\|EnsemblGenome=GSU2836\|UniProtKB=Q749A8	Geobacter sulfurreducens	O
BRADU\|Gene=adk\|UniProtKB=Q89JA5	Bradyrhizobium diazoefficiens	O
COXBU\|EnsemblGenome=CBU_0454\|UniProtKB=Q83E75	Coxiella burnetii	O
HAEIN\|EnsemblGenome=HI_0349\|UniProtKB=P24323	Haemophilus influenzae	O
SHEON\|Gene_OrderedLocusName=SO_2018\|UniProtKB=Q8EFF5	Shewanella oneidensis	O
VIBCH\|EnsemblGenome=VC_0986\|UniProtKB=Q9KTB7	Vibrio cholerae	O
PSEAE\|EnsemblGenome=PA3686\|UniProtKB=Q9HXV4	Pseudomonas aeruginosa	O
SALTY\|EnsemblGenome=STM0488\|UniProtKB=P0A1V4	Salmonella typhimurium	O
YERPE\|EnsemblGenome=YP_0812\|UniProtKB=O69172	Yersinia pestis	O
ECOLI\|EnsemblGenome=BAE76253\|UniProtKB=P69441	Escherichia coli	O