PANTHER Gene Information

Gene Symbol(s):

CHLRE_07g351050v5

Organism:

Chlamydomonas reinhardtii

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Tree

Gene Name:

Uncharacterized protein

Gene ID:

CHLRE_07g351050v5

Protein ID:

A0A2K3DLC5

Persistent Id:

PTN004628092

Alternate Ids:

A0A2K3DLC5(Synonym)	CM008968(EMBL)
CHLRE_07g351050v5(Ensembl)	PNW81323(EMBL-CDS)
A0A2K3DLC5_CHLRE(UniProtKB-ID)
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PANTHER Classification

PANTHER Family:	TUMOR SUSCEPTIBILITY GENE 101 PROTEIN-RELATED (PTHR23306)
PANTHER Subfamily:	TUMOR SUPPRESSOR PROTEIN 101 (PTHR23306:SF3)
PANTHER GO-slim Molecular Function:	ubiquitin binding
PANTHER GO-slim Biological Process:	endosome to lysosome transport
PANTHER GO-slim Cellular Component:	ESCRT I complex
PANTHER protein class:	ubiquitin-protein ligase

Pathway Categories:

No pathway information available

GENE ONTOLOGY DATABASE ANNOTATIONS

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GO MF Complete:	ubiquitin binding
GO BP Complete:	protein modification process, protein transport, endosome to lysosome transport
GO CC Complete:	endosome, ESCRT I complex

REACTOME DATABASE ANNOTATIONS Click to Expand...

Reactome Pathways:

Orthologs

ID	Organism	Type
HUMAN\|HGNC=30866\|UniProtKB=Q8IX04	Homo sapiens	O
PANTR\|Ensembl=ENSPTRG00000003425.7\|UniProtKB=H2RCW9	Pan troglodytes	O
GORGO\|Ensembl=ENSGGOG00000010367.3\|UniProtKB=G3R4C1	Gorilla gorilla gorilla	O
MACMU\|Ensembl=ENSMMUG00000000336.4\|UniProtKB=H9FWP8	Macaca mulatta	O
MOUSE\|MGI=MGI=1860490\|UniProtKB=Q3U1V6	Mus musculus	O
RAT\|RGD=1587416\|UniProtKB=A0A8I6ALM7	Rattus norvegicus	O
BOVIN\|Ensembl=ENSBTAG00000013568.5\|UniProtKB=A5PJF9	Bos taurus	O
PIG\|Ensembl=ENSSSCG00005016101.1\|UniProtKB=A0A4X1T5Q2	Sus scrofa	O
FELCA\|Ensembl=ENSFCAG00000015742.6\|UniProtKB=M3WN71	Felis catus	O
CANLF\|Ensembl=ENSCAFG00845017016.1\|UniProtKB=A0A8I3P1P9	Canis lupus familiaris	O
MONDO\|Ensembl=ENSMODG00000006389.4\|UniProtKB=A0A5F8GVK3	Monodelphis domestica	O
ORNAN\|Ensembl=ENSOANG00000050347.1\|UniProtKB=A0A6I8PDA4	Ornithorhynchus anatinus	O
CHICK\|Ensembl=ENSGALG00010024496.1\|UniProtKB=A0A8V1A5C6	Gallus gallus	O
ANOCA\|Ensembl=ENSACAG00000000063.4\|UniProtKB=G1K890	Anolis carolinensis	O
DANRE\|ZFIN=ZDB-GENE-040801-138\|UniProtKB=Q6DBY5	Danio rerio	O
HUMAN\|HGNC=15971\|UniProtKB=Q99816	Homo sapiens	LDO
PANTR\|Ensembl=ENSPTRG00000003424.6\|UniProtKB=A0A2I3SDN5	Pan troglodytes	LDO
GORGO\|Ensembl=ENSGGOG00000010359.3\|UniProtKB=G3R4B1	Gorilla gorilla gorilla	LDO
MACMU\|Ensembl=ENSMMUG00000014613.4\|UniProtKB=G7NDS4	Macaca mulatta	LDO
MOUSE\|MGI=MGI=106581\|UniProtKB=Q61187	Mus musculus	LDO
RAT\|RGD=3909\|UniProtKB=Q6IRE4	Rattus norvegicus	LDO
BOVIN\|Ensembl=ENSBTAG00000013563.6\|UniProtKB=A3KN51	Bos taurus	LDO
HORSE\|Ensembl=ENSECAG00000004362.4\|UniProtKB=A0A9L0RHA1	Equus caballus	LDO
FELCA\|Ensembl=ENSFCAG00000028742.4\|UniProtKB=A0A337SU32	Felis catus	LDO
CANLF\|Ensembl=ENSCAFG00845016679.1\|UniProtKB=A0A8I3P8P4	Canis lupus familiaris	LDO
MONDO\|Ensembl=ENSMODG00000006410.3\|UniProtKB=F7FD96	Monodelphis domestica	LDO
ORNAN\|Ensembl=ENSOANG00000008368.3\|UniProtKB=A0A6I8PK40	Ornithorhynchus anatinus	LDO
ANOCA\|Ensembl=ENSACAG00000000062.4\|UniProtKB=A0A803TR58	Anolis carolinensis	LDO
XENLA\|Gene=tsg101.L\|UniProtKB=A0A8J1MW83	Xenopus laevis	O
XENLA\|Gene=MGC81040\|UniProtKB=Q6NUD5	Xenopus laevis	LDO
XENTR\|Gene=tsg101\|UniProtKB=A0A8J1JD66	Xenopus tropicalis	LDO
ORYLA\|Ensembl=ENSORLG00000023170.1\|UniProtKB=A0A3B3HBA2	Oryzias latipes	O
DANRE\|ZFIN=ZDB-GENE-051127-21\|UniProtKB=Q2YDS7	Danio rerio	O
ORYLA\|Ensembl=ENSORLG00000030185.1\|UniProtKB=A0A3B3IPA0	Oryzias latipes	LDO
DANRE\|ZFIN=ZDB-GENE-130530-542\|UniProtKB=E7F4Y8	Danio rerio	O
DANRE\|ZFIN=ZDB-GENE-030217-1\|UniProtKB=Q6IQ70	Danio rerio	LDO
LEPOC\|Ensembl=ENSLOCG00000000710.1\|UniProtKB=W5LXD4	lepisosteus oculatus	LDO
CIOIN\|Ensembl=ENSCING00000021102.1\|UniProtKB=H2XP92	Ciona intestinalis	LDO
BRAFL\|Gene=LOC118406970\|UniProtKB=A0A9J7KK24	Branchiostoma floridae	O
BRAFL\|Gene=LOC118421770\|UniProtKB=A0A9J7LLM7	Branchiostoma floridae	O
BRAFL\|Gene=LOC118406656\|UniProtKB=A0A9J7HNB8	Branchiostoma floridae	O
BRAFL\|Gene=LOC118406596\|UniProtKB=A0A9J7HNJ2	Branchiostoma floridae	LDO
STRPU\|EnsemblGenome=LOC105439760\|UniProtKB=A0A7M7P4V5	Strongylocentrotus purpuratus	LDO
STRPU\|EnsemblGenome=LOC763823\|UniProtKB=A0A7M7SZX9	Strongylocentrotus purpuratus	LDO
CAEEL\|WormBase=WBGene00015658\|UniProtKB=O76258	Caenorhabditis elegans	LDO
DROME\|FlyBase=FBgn0036666\|UniProtKB=Q9VVA7	Drosophila melanogaster	LDO
ANOGA\|EnsemblGenome=AGAP005934\|UniProtKB=Q7Q6B6	Anopheles gambiae	LDO
TRICA\|EnsemblGenome=TC002425\|UniProtKB=D6WIH5	Tribolium castaneum	LDO
DAPPU\|Gene_ORFName=DAPPUDRAFT_65881\|UniProtKB=E9HTS3	Daphnia pulex	LDO
IXOSC\|EnsemblGenome=ISCW014292\|UniProtKB=B7QIP7	Ixodes scapularis	LDO
HELRO\|EnsemblGenome=HelroG185660\|UniProtKB=T1FN37	helobdella robusta	LDO
NEMVE\|EnsemblGenome=NEMVEDRAFT_v1g244032\|UniProtKB=A7SAT5	Nematostella vectensis	LDO
TRIAD\|EnsemblGenome=TriadG33034\|UniProtKB=B3SBY4	Trichoplax adhaerens	LDO
YEAST\|SGD=S000000514\|UniProtKB=P25604	Saccharomyces cerevisiae	LDO
EREGS\|EnsemblGenome=AGOS_AAL006C\|UniProtKB=Q75EU1	Eremothecium gossypii	LDO
CANAL\|CGD=CAL0000187259\|UniProtKB=Q59WF8	Candida albicans	LDO
EMENI\|EnsemblGenome=ANIA_02521\|UniProtKB=Q5BAA9	Emericella nidulans	LDO
ASPFU\|EnsemblGenome=AFUA_3G14380\|UniProtKB=Q4WYV3	Neosartorya fumigata	LDO
NEUCR\|EnsemblGenome=NCU05753\|UniProtKB=Q7S4R9	Neurospora crassa	LDO
SCLS1\|EnsemblGenome=SS1G_10308\|UniProtKB=A7EY93	Sclerotinia sclerotiorum	LDO
PHANO\|EnsemblGenome=SNOG_16164\|UniProtKB=Q0TWM8	Phaeosphaeria nodorum	LDO
CRYNJ\|EnsemblGenome=CNC02240\|UniProtKB=Q5KKQ1	Cryptococcus neoformans	LDO
USTMA\|EnsemblGenome=UMAG_11474\|UniProtKB=A0A0D1E2K5	Ustilago maydis	LDO
PUCGT\|EnsemblGenome=PGTG_14838\|UniProtKB=E3KWF8	Puccinia graminis	LDO
BATDJ\|Gene_ORFName=BATDEDRAFT_90872\|UniProtKB=F4P9F9	Batrachochytrium dendrobatidis	LDO
ENTH1\|EnsemblGenome=CL6EHI_194210\|UniProtKB=C4LZT6	Entamoeba histolytica	LDO
ENTH1\|EnsemblGenome=CL6EHI_131530\|UniProtKB=C4M9Z1	Entamoeba histolytica	O
ENTH1\|EnsemblGenome=CL6EHI_193530\|UniProtKB=B1N3Q7	Entamoeba histolytica	O
ENTH1\|EnsemblGenome=CL6EHI_178530\|UniProtKB=C4LUR9	Entamoeba histolytica	O
ENTH1\|EnsemblGenome=CL6EHI_135460\|UniProtKB=C4M9S4	Entamoeba histolytica	O
DICDI\|dictyBase=DDB_G0286797\|UniProtKB=Q54LJ3	Dictyostelium discoideum	LDO
DICPU\|EnsemblGenome=DICPUDRAFT_155276\|UniProtKB=F0ZTJ6	Dictyostelium purpureum	LDO
LEIMA\|EnsemblGenome=LMJF_32_0985\|UniProtKB=E9AEF7	Leishmania major	LDO
TRYB2\|Gene_ORFName=Tb11.01.3960\|UniProtKB=Q382W6	Trypanosoma brucei	O
TRYB2\|Gene_ORFName=Tb11.01.5840\|UniProtKB=Q382B4	Trypanosoma brucei	LDO
TRIV3\|Gene_ORFName=TVAG_353480\|UniProtKB=A2EN70	Trichomonas vaginalis	LDO
TRIV3\|Gene_ORFName=TVAG_193190\|UniProtKB=A2DH33	Trichomonas vaginalis	O
TRIV3\|Gene_ORFName=TVAG_128690\|UniProtKB=A2E4C0	Trichomonas vaginalis	O
PARTE\|EnsemblGenome=GSPATT00005706001\|UniProtKB=A0BV93	Paramecium tetraurelia	O
PARTE\|EnsemblGenome=GSPATT00009545001\|UniProtKB=A0CR15	Paramecium tetraurelia	LDO
THAPS\|EnsemblGenome=THAPS_23630\|UniProtKB=B5YNX5	Thalassiosira pseudonana	LDO
PHYRM\|Gene=H3GRU5_PHYRM\|UniProtKB=H3GRU5	Phytophthora ramorum	LDO
AMBTC\|EnsemblGenome=AMTR_s00002p00269040\|UniProtKB=W1P3D6	Amborella trichopoda	O
AMBTC\|EnsemblGenome=AMTR_s00051p00167090\|UniProtKB=U5D8G3	Amborella trichopoda	O
AMBTC\|EnsemblGenome=AMTR_s00051p00166200\|UniProtKB=U5D5G4	Amborella trichopoda	LDO
ZOSMR\|Gene_ORFName=ZOSMA_94G00270\|UniProtKB=A0A0K9NIP5	Zostera marina	O
MUSAM\|EnsemblGenome=Ma03_g32210\|UniProtKB=A0A804IIP9	Musa acuminata subsp. malaccensis	O
MUSAM\|EnsemblGenome=Ma04_g14860\|UniProtKB=A0A804IPV1	Musa acuminata subsp. malaccensis	O
SETIT\|EnsemblGenome=SETIT_024977mg\|UniProtKB=A0A0Q3TSZ9	Setaria italica	O
SORBI\|EnsemblGenome=SORBI_3008G088803\|UniProtKB=A0A109NDP8	Sorghum bicolor	O
MAIZE\|EnsemblGenome=Zm00001eb409970\|UniProtKB=A0A804RAY1	Zea mays	O
NELNU\|Gene_ORFName=HUJ06_007277\|UniProtKB=A0A1U8BD99	Nelumbo nucifera	O
NELNU\|Gene_ORFName=HUJ06_021168\|UniProtKB=A0A1U8BPA6	Nelumbo nucifera	O
NELNU\|Gene_ORFName=HUJ06_007278\|UniProtKB=A0A1U8B0X3	Nelumbo nucifera	O
HELAN\|EnsemblGenome=HanXRQr2_Chr10g0423261\|UniProtKB=A0A251TIQ9	Helianthus annuus	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a020747mg\|UniProtKB=A0A022S1A0	Erythranthe guttata	O
CAPAN\|EnsemblGenome=T459_08838\|UniProtKB=A0A2G2ZXL5	Capsicum annuum	O
SOLTU\|EnsemblGenome=PGSC0003DMG400019336\|UniProtKB=M1BPA9	Solanum tuberosum	O
SOLLC\|EnsemblGenome=Solyc09g065770.1\|UniProtKB=A0A3Q7I4F2	Solanum lycopersicum	O
TOBAC\|Gene=LOC107826539\|UniProtKB=A0A1S4D6J9	Nicotiana tabacum	O
TOBAC\|Gene=LOC107788448\|UniProtKB=A0A1S3ZMX8	Nicotiana tabacum	O
VITVI\|EnsemblGenome=Vitvi07g00060\|UniProtKB=F6HBJ5	Vitis vinifera	O
EUCGR\|EnsemblGenome=EUGRSUZ_B01481\|UniProtKB=A0A059D226	Eucalyptus grandis	O
BRANA\|Gene_ORFName=GSBRNA2T00005351001\|UniProtKB=A0A078FYV7	Brassica napus	O
BRACM\|EnsemblGenome=Bra017097\|UniProtKB=M4DKR6	Brassica campestris	O
ARATH\|TAIR=locus=2040394\|UniProtKB=F4ITY4	Arabidopsis thaliana	O
GOSHI\|Gene=LOC107955479\|UniProtKB=A0A1U8P6A7	Gossypium hirsutum	O
GOSHI\|Gene=LOC107941651\|UniProtKB=A0A1U8MV58	Gossypium hirsutum	O
THECC\|EnsemblGenome=TCM_004230\|UniProtKB=A0A061DQB8	Theobroma cacao	O
CITSI\|Gene_ORFName=CISIN_1g047127mg\|UniProtKB=A0A067FYR1	Citrus sinensis	O
POPTR\|EnsemblGenome=POPTR_002G046200v3\|UniProtKB=A0A2K2BDE4	Populus trichocarpa	O
MANES\|EnsemblGenome=MANES_05G083600\|UniProtKB=A0A2C9VWV7	Manihot esculenta	O
RICCO\|Gene_ORFName=RCOM_0991540\|UniProtKB=B9S4R4	Ricinus communis	O
ZOSMR\|Gene_ORFName=ZOSMA_215G00100\|UniProtKB=A0A0K9PK70	Zostera marina	LDO
MUSAM\|EnsemblGenome=Ma03_g28060\|UniProtKB=A0A804IH61	Musa acuminata subsp. malaccensis	O
MUSAM\|EnsemblGenome=Ma10_g02590\|UniProtKB=A0A804KRU1	Musa acuminata subsp. malaccensis	O
MUSAM\|EnsemblGenome=Ma11_g06970\|UniProtKB=A0A804L544	Musa acuminata subsp. malaccensis	LDO
MAIZE\|EnsemblGenome=Zm00001eb259630\|UniProtKB=A0A804PQD0	Zea mays	LDO
ORYSJ\|EnsemblGenome=Os02g0833300\|UniProtKB=Q6ESB7	Oryza sativa	LDO
BRADI\|EnsemblGenome=BRADI_1g04460v3\|UniProtKB=A0A0Q3GQ74	Brachypodium distachyon	O
HORVV\|EnsemblGenome=HORVU.MOREX.r3.5HG0523850\|UniProtKB=A0A287SQ79	Hordeum vulgare subsp. vulgare	O
WHEAT\|EnsemblGenome=TraesCS5A02G463000\|UniProtKB=A0A3B6KS41	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS5D02G475800\|UniProtKB=A0A3B6MZ42	Triticum aestivum	O
BRADI\|EnsemblGenome=BRADI_3g60837v3\|UniProtKB=A0A0Q3I8R7	Brachypodium distachyon	LDO
HORVV\|EnsemblGenome=HORVU.MOREX.r3.5HG0504530\|UniProtKB=M0XLK7	Hordeum vulgare subsp. vulgare	LDO
WHEAT\|EnsemblGenome=TraesCS5D02G365300\|UniProtKB=A0A3B6MWR5	Triticum aestivum	LDO
WHEAT\|EnsemblGenome=TraesCS5B02G358900\|UniProtKB=A0A3B6LS51	Triticum aestivum	O
WHEAT\|EnsemblGenome=TraesCS5A02G356400\|UniProtKB=A0A3B6KKU0	Triticum aestivum	LDO
NELNU\|Gene=LOC104609678\|UniProtKB=A0A1U8B0V4	Nelumbo nucifera	LDO
NELNU\|Gene=LOC104601457\|UniProtKB=A0A1U8A8W4	Nelumbo nucifera	O
SPIOL\|Gene=LOC110794291\|UniProtKB=A0A9R0ITM2	Spinacia oleracea	LDO
HELAN\|EnsemblGenome=HanXRQr2_Chr04g0175941\|UniProtKB=A0A251VU42	Helianthus annuus	LDO
LACSA\|Gene_ORFName=LSAT_V11C700347150\|UniProtKB=A0A9R1X546	Lactuca sativa	LDO
HELAN\|EnsemblGenome=HanXRQr2_Chr17g0819101\|UniProtKB=A0A251T572	Helianthus annuus	O
LACSA\|Gene_ORFName=LSAT_V11C400172490\|UniProtKB=A0A9R1XFA3	Lactuca sativa	O
ERYGU\|Gene_ORFName=MIMGU_mgv1a007629mg\|UniProtKB=A0A022Q0B4	Erythranthe guttata	LDO
SOLTU\|EnsemblGenome=PGSC0003DMG400002673\|UniProtKB=M0ZS40	Solanum tuberosum	O
SOLLC\|EnsemblGenome=Solyc09g009270.2\|UniProtKB=A0A3Q7HZ13	Solanum lycopersicum	O
TOBAC\|Gene=LOC107786494\|UniProtKB=A0A1S3ZGJ7	Nicotiana tabacum	O
TOBAC\|Gene=LOC107767728\|UniProtKB=A0A1S3XQZ1	Nicotiana tabacum	O
CAPAN\|EnsemblGenome=T459_25806\|UniProtKB=A0A2G2YLT3	Capsicum annuum	LDO
SOLTU\|EnsemblGenome=PGSC0003DMG400008386\|UniProtKB=M1AFF3	Solanum tuberosum	LDO
SOLLC\|EnsemblGenome=Solyc10g085590.1\|UniProtKB=A0A3Q7JGU6	Solanum lycopersicum	LDO
TOBAC\|Gene=LOC107791870\|UniProtKB=A0A1S3ZYW5	Nicotiana tabacum	LDO
TOBAC\|Gene=LOC107828341\|UniProtKB=A0A1S4DCE8	Nicotiana tabacum	LDO
VITVI\|EnsemblGenome=Vitvi15g00653\|UniProtKB=F6GV51	Vitis vinifera	O
VITVI\|EnsemblGenome=Vitvi05g01184\|UniProtKB=F6GWC1	Vitis vinifera	LDO
BRACM\|EnsemblGenome=Bra023439\|UniProtKB=M4E3T6	Brassica campestris	LDO
ARATH\|TAIR=locus=2159113\|UniProtKB=Q9FFY6	Arabidopsis thaliana	LDO
BRANA\|Gene_ORFName=DARMORV10_C02P33210.1\|UniProtKB=A0A078G4R5	Brassica napus	O
BRANA\|Gene_ORFName=DARMORV10_A02P20570.1\|UniProtKB=A0A078I9C5	Brassica napus	O
BRACM\|EnsemblGenome=Bra008248\|UniProtKB=M4CVK1	Brassica campestris	O
BRACM\|EnsemblGenome=Bra001451\|UniProtKB=M4CB71	Brassica campestris	O
ARATH\|TAIR=locus=2092477\|UniProtKB=Q9LHG8	Arabidopsis thaliana	O
GOSHI\|Gene=LOC107955341\|UniProtKB=A0A1U8P639	Gossypium hirsutum	LDO
THECC\|EnsemblGenome=TCM_044692\|UniProtKB=A0A061FR54	Theobroma cacao	LDO
CITSI\|Gene_ORFName=CISIN_1g039489mg\|UniProtKB=A0A067DDL2	Citrus sinensis	LDO
POPTR\|EnsemblGenome=POPTR_003G191100v3\|UniProtKB=B9GXA7	Populus trichocarpa	O
POPTR\|EnsemblGenome=POPTR_001G034200v3\|UniProtKB=A0A2K2BRT6	Populus trichocarpa	LDO
MANES\|EnsemblGenome=MANES_04G154600\|UniProtKB=A0A2C9W5C3	Manihot esculenta	O
MANES\|EnsemblGenome=MANES_11G009300\|UniProtKB=A0A2C9UXE2	Manihot esculenta	LDO
RICCO\|Gene_ORFName=RCOM_0621510\|UniProtKB=B9SHU4	Ricinus communis	LDO
SOYBN\|EnsemblGenome=GLYMA_12G025300\|UniProtKB=I1LPI0	Glycine max	O
SOYBN\|EnsemblGenome=GLYMA_11G099100\|UniProtKB=I1LIR0	Glycine max	LDO
MEDTR\|EnsemblGenome=MTR_4g075250\|UniProtKB=G7JPJ7	Medicago truncatula	LDO
JUGRE\|Gene=LOC108989390\|UniProtKB=A0A2I4EGI7	Juglans regia	O
JUGRE\|Gene=LOC118344520\|UniProtKB=A0A6P9E011	Juglans regia	LDO
JUGRE\|EnsemblGenome=Jr14_02310\|UniProtKB=A0A2I4GZN3	Juglans regia	LDO
CUCSA\|EnsemblGenome=Csa_5G583320\|UniProtKB=A0A0A0KT64	Cucumis sativus	LDO
PRUPE\|EnsemblGenome=PRUPE_2G048600\|UniProtKB=M5WZN4	Prunus persica	LDO
SELML\|EnsemblGenome=SELMODRAFT_138144\|UniProtKB=D8TEQ5	Selaginella moellendorffii	LDO
SELML\|EnsemblGenome=SELMODRAFT_129226\|UniProtKB=D8T0J2	Selaginella moellendorffii	LDO
PHYPA\|EnsemblGenome=Pp3c27_3040\|UniProtKB=A0A2K1IAG6	Physcomitrella patens	O
PHYPA\|EnsemblGenome=Pp3c16_2240\|UniProtKB=A9TJ74	Physcomitrella patens	LDO
MARPO\|EnsemblGenome=MARPO_0055s0078\|UniProtKB=A0A2R6WVC7	Marchantia polymorpha	O
MARPO\|EnsemblGenome=MARPO_0030s0083\|UniProtKB=A0A2R6X8B5	Marchantia polymorpha	LDO
KLENI\|Gene_ORFName=KFL_003310130\|UniProtKB=A0A1Y1IDC2	Klebsormidium nitens	O
KLENI\|Gene_ORFName=KFL_000990220\|UniProtKB=A0A1Y1HW97	Klebsormidium nitens	LDO